A group of proteins required for telomere positioning. (A) The Bqt3–Bqt4 complex provides an attachment surface for telomeres to the NE in vegetative cells. (B and C) The Bqt1–Bqt2 complex binds to Rap1 at the telomere upon mating pheromone signaling during meiosis, followed by accumulation of the Sad1–Kms1 complex to the telomere through interaction with Bqt1.