Alternative mechanisms for chromosome segregation in mammalian mitosis. Chromosomes that fail to establish direct connections with the spindle poles nevertheless maintain their position near the spindle equator and segregate during anaphase via dynein-mediated force that acts upon a short K-fiber attached to the kinetochore. Poleward movements of these chromosomes are not coupled with depolymerization of K-fiber MTs at the kinetochore (Pacman mechanism). Instead the entire short fiber with the attached kinetochore is transported toward the spindle pole.