Table 1.

PKCδ deficiency reported in the literature

ReferenceAge at onsetSexEthnicityGenetic variantInitial presentationInfectionsLymphoproliferationAutoimmunityLaboratoryTreatment(s)Age at last follow-upAlive/dead
Salzer et al., 2013, PT1 (2) Infancy Turkish c.1352+1G>A Recurrent infections URTI (i.e., pneumonia), LRTI, UTIs, gastroenteritis otitis media, and herpes virus viremia Hepatosplenomegaly and lymphadenopathy Nephrotic syndrome, nonspecific reactive follicular hyperplasia, relapsing polychondritis, hypothyroidism, antiphospholipid syndrome, and SLE Progressive reduction in B cells, reduced memory B cells, increased CD21low B cells, low IgG levels, and elevated IgA and IgM levels Steroids, rituximab, MMF, IVIG, enalapril, anticoagulants, and thyroid hormone replacement 12 years Alive 
Belot et al., 2013, PTV1, V3, V4 (10) 10 years White, of Northern European extraction c.1528G>A Discoid lupus rash and arthritis None reported Hepatomegaly Alopecia, lupus nephritis, arthritis, chronic cutaneous lupus, WHO type 2 glomerulonephritis with severe nephrotic syndrome, chronic renal failure, SLE, and CNS vasculitis Lymphopenia (B and T cells), positive ANAs (1:640) and anti-dsDNA antibodies, low C3 and C4, and elevated IgA Steroids, hydroxychloroquine,
Azathioprine, methotrexate, and
MMF 
24 years Alive 
3 years White, of Northern European extraction c.1528G>A Diffuse lymphoproliferation and AIHA P. aeruginosa septic shock Lymphadenopathy, multiple compressive adenopathy with mediastinal involvement and hepatosplenomegaly SLE, AIHA, serositis, pancreatitis, ITP, lupus rash, and lupus nephritis Lymphopenia, but increased CD3+ T cells, undetectable double-negative TCRα/β T cells, positive ANAs (1:640) and anti-dsDNA antibodies, and low C3 and C4 Immunosuppressive treatments 8 years Died at age 13 years (end-stage kidney failure complicated by Pseudomonas sustained septic shock) 
6 years White, of Northern European extraction c.1528G>A Lupus nephritis ​ None Renal flare with malar rash and arthritis SLE Increased immature and naive B cells, reduced memory B cells, positive ANAs, and anti-dsDNA antibodies Steroids (prednisolone) and
MMF 
13 years Alive 
Kuehn et al., 2013, PT1 (1) 3 years Hispanic c.1840C>T Recurrent otitis, sinusitis, persistent lymphadenopathy, hepatosplenomegaly, and intermittent fevers Recurrent otitis media/sinusitis, and persistent EBV Hepatosplenomegaly and mediastinal lymphadenitis (with superior vena cava syndrome) SLE and AIHA CD5+ majority B lymphocytosis; reduced class-switched memory B cells; elevated DNT αβ and DNT γδ; low levels of NK cells and NK cell cytolytic activity; increased ESR, CRP, ALT, and AST; hypergammaglobulinemia; positive ANAs; anti-RNP; anti-Smith; anti-SSA Steroids (prednisone) and sirolimus 7 years Alive 
Kiykim et al., 2015, PT6 (11) Infancy Turkish c.742G>A Recurrent fever Pneumonia, gastroenteritis, and CMV Hepatosplenomegaly and lymphadenopathy SLE Naive majority B lymphocytosis and CD21low, reduced levels of switched memory B cells, increased IgM, and reduced NK functionality Ganciclovir, IVIG, topical steroids, hydroxychloroquine,
IVIG, and prophylactic antibiotics 
3.5 years Alive 
Lei et al., 2018, PT II-1, PT II-2, and PT 11-3 (12) 1 year Endogamous Pakistani c.1294G>T Constitutional symptoms (intermittent fever, night sweats, fatigue), severe thrombocytopenia, photosensitive and petechial rash, scarring alopecia, and hepatosplenomegaly NR Splenomegaly and lymphadenopathy SLE (acuta cutaneous lupus, oral/nasal ulcers) AIHA, thrombocytopenia, leukopenia, positive ANA (1:1,280), anti-dsDNA, antiphospholipid antibody, low complement, elevated CRP and ESR, elevated IgG, and low levels of C4 Steroid, rituximab (discontinued for adverse reaction),
IVIG, hydroxychloroquine,
Azathioprine, and
Ofatumumab 
NR Alive 
1 year Endogamous Pakistani c.1294G>T Constitutional symptoms (intermittent fever, night sweats, fatigue), severe thrombocytopenia, photosensitive and petechial rash, scarring alopecia, hepatosplenomegaly NR Hepatosplenomegaly, lymphadenopathy SLE (acute cutaneous lupus, serositis, and renal impairment) AIHA, thrombocytopenia, leukopenia, slight neutropenia, positive ANA (1:1,280), anti-dsDNA, antiphospholipid antibody, and low complement, elevated CRP and ESR, elevated albumin/creatinine ratio, hypergammaglobulinemia, and low levels of C4 Steroid,
IVIG, rituximab (discontinued for adverse reaction),h
ydroxychloroquine,
Azathioprine, and
Ofatumumab 
NR Alive 
26 mo Endogamous Pakistani c.1294G>T Constitutional symptoms (intermittent fever, night sweats, fatigue), severe thrombocytopenia, photosensitive and petechial rash, scarring alopecia, and hepatosplenomegaly NR Hepatosplenomegaly SLE (acute cutaneous lupus) AIHA, thrombocytopenia, positive ANA (1:1,280), anti-dsDNA, anti-Smith antibody, and low complement Steroid (IV methylprednisolone),
IVIG, and rituximab 
NR Alive 
Sharifinejad et al., 2020, PT10 (13) Infancy Afghan c.1293_1294insA Recurrent oral candidiasis, nonsevere upper respiratory and gastrointestinal tract infections, and food allergy Pneumonia, oral candidiasis, and recurrent gastroenteritis Hepatosplenomegaly and lymphadenopathy SLE Increased DNT, reduced NK number, increased IgG, and nonprotective antidiphtheria Corticosteroids, antibiotic prophylaxis 13 years Alive 
Neehus et al., 2021, PT13-17 (14) 1 year Turkish c.571+2dup BCGosis, fever, left axillary, retroauricular, and cervical lymphadenopathies BCGosis, Salmonella bacteremia, Staphylococcus aureus lymphadenitis, and recurrent gingivitis Lymphadenopathy and hepatosplenomegaly SLE Elevated naive B cells and reduced B memory, with elevated CD21low and transitional cells, hypogammaglobulinemia, ANA+, and anti-dsDNA+ Steroids
IVIG
Antimycobacterial (ceftriaxone); antibiotic prophylaxis (amoxicillin) 
14 years Alive 
7 mo Turkish c.1384C>T Fever, diarrhea, hepatosplenomegaly, failure to thrive, left axillary lymphadenopathy with purulent discharge, and BCGitis BCGitis, recurrent gastroenteritis Hepatosplenomegaly AIHA Impaired DHR test and ANA+ Steroids
Isoniazid
Rifampicin 
10 years Alive 
7 mo Turkish c.1384C>T BCGitis BCGitis, recurrent gastroenteritis and shingles Lymphadenopathy and hepatosplenomegaly AIHA Impaired DHR test and ANA+ Steroids
Isoniazid
Rifampicin
IVIG 
7 years Alive 
1 year Irani c.642del BCGitis BCGitis Hepatosplenomegaly and lymphadenopathy None Impaired DHR test and nonprotective antibody responses to tetanus and diphtheria Isoniazid
Rifampicin Azithromycin 
3 years Alive 
NR Irani c.642del Asymptomatic None None None Impaired DHR test No treatment 6 years Alive 
Gu et al., 2021, PT16-17 (15) and Jefferson et al., 2023 (3) 6 y Chinese c.36T>G Lymphadenopathy, splenomegaly, and recurrent infections NR Splenomegaly and lymphadenopathy Pancytopenia Positive antibodies for EBV, CMV, herpes simplex virus, and rubella viruses, increased TCRαβ DNT, increased CD8+ T cells and IgG, and low levels of NK cells Steroids, IRT, and sirolimus 10 years Alive 
NR Chinese c.36T>G Lymphadenopathy, splenomegaly, and anemia NR Splenomegaly and lymphadenopathy Anemia NR NR NR NR 
Yang et al., 2022, PT18 (18) 5 mo Chinese c.661C>T Recurrent fever, respiratory tract infections, and lymphadenopathy URTIs and
BCGosis 
Multiple lymphadenopathy (abdominal mesenteric lymph nodes invading the abdominal wall), bone marrow hyperplasia, and hepatosplenomegaly NR Increased DNT, elevated CD8+ T cells, low levels of CD4+ T cells, and CRP positive Isoniazid, rifampicin, ethambutol, sirolimus,
IFN-γ, and antibiotics 
8 years Alive 
Neehus et al., 2022, PT19 (19) 2 years American c.285C>A/c.376C>T URTI, lymphadenitis Burkholderia cepacia and Mycobacterium lentiflavum lymphadenopathy Infectious lymphadenopathy None Low levels of CD4+ T cells, decreased relative proportions of CD27+IgD+ and CD27+IgD memory B cells, and abnormally low DHR test Cotrimoxazole, isoniazid, and
Rifampicin 
5 years Alive 
Roderick et al., 2023, PT20-21 (9) 9 mo British c.788-2A>G/c.571C>T Listeria meningitis Listeria meningitis, Enterococcus faecium bacteremia, varicella and shingles, invasive Candida infection, recurrent episodes of Achromobacter xylosoxidans cervical lymphadenitis, and early-onset enterocolitis Lymphoid hyperplasia and splenomegaly SLE, AIHA, and ITP Increased DNTs, low B cells with low IgG with high IgM, positive anticardiolipin antibodies, Coombs positive, and positive thyroid peroxidase antibodies Steroids,
IVIG, rituximab,
Sirolimus,
Hydroxychloroquine,
HSCT, and cyclosporine (for GVHD prophylaxis) 
NR Alive 
20 mo British c.788-2A>G/c.571C>T Sepsis due to nontypeable Haemophilus influenzae Early-onset enterocolitis Lymphoid hyperplasia, splenomegaly SLE Increased DNTs, low B cells with low IgG and high IgM, positive anticardiolipin antibodies, and GAD antibody-positive IVIG replacement therapy, rituximab, sirolimus (stopped because of oral ulceration),
MMF, and hydroxychloroquine
HSCT 
8.5 years Alive 
ReferenceAge at onsetSexEthnicityGenetic variantInitial presentationInfectionsLymphoproliferationAutoimmunityLaboratoryTreatment(s)Age at last follow-upAlive/dead
Salzer et al., 2013, PT1 (2) Infancy Turkish c.1352+1G>A Recurrent infections URTI (i.e., pneumonia), LRTI, UTIs, gastroenteritis otitis media, and herpes virus viremia Hepatosplenomegaly and lymphadenopathy Nephrotic syndrome, nonspecific reactive follicular hyperplasia, relapsing polychondritis, hypothyroidism, antiphospholipid syndrome, and SLE Progressive reduction in B cells, reduced memory B cells, increased CD21low B cells, low IgG levels, and elevated IgA and IgM levels Steroids, rituximab, MMF, IVIG, enalapril, anticoagulants, and thyroid hormone replacement 12 years Alive 
Belot et al., 2013, PTV1, V3, V4 (10) 10 years White, of Northern European extraction c.1528G>A Discoid lupus rash and arthritis None reported Hepatomegaly Alopecia, lupus nephritis, arthritis, chronic cutaneous lupus, WHO type 2 glomerulonephritis with severe nephrotic syndrome, chronic renal failure, SLE, and CNS vasculitis Lymphopenia (B and T cells), positive ANAs (1:640) and anti-dsDNA antibodies, low C3 and C4, and elevated IgA Steroids, hydroxychloroquine,
Azathioprine, methotrexate, and
MMF 
24 years Alive 
3 years White, of Northern European extraction c.1528G>A Diffuse lymphoproliferation and AIHA P. aeruginosa septic shock Lymphadenopathy, multiple compressive adenopathy with mediastinal involvement and hepatosplenomegaly SLE, AIHA, serositis, pancreatitis, ITP, lupus rash, and lupus nephritis Lymphopenia, but increased CD3+ T cells, undetectable double-negative TCRα/β T cells, positive ANAs (1:640) and anti-dsDNA antibodies, and low C3 and C4 Immunosuppressive treatments 8 years Died at age 13 years (end-stage kidney failure complicated by Pseudomonas sustained septic shock) 
6 years White, of Northern European extraction c.1528G>A Lupus nephritis ​ None Renal flare with malar rash and arthritis SLE Increased immature and naive B cells, reduced memory B cells, positive ANAs, and anti-dsDNA antibodies Steroids (prednisolone) and
MMF 
13 years Alive 
Kuehn et al., 2013, PT1 (1) 3 years Hispanic c.1840C>T Recurrent otitis, sinusitis, persistent lymphadenopathy, hepatosplenomegaly, and intermittent fevers Recurrent otitis media/sinusitis, and persistent EBV Hepatosplenomegaly and mediastinal lymphadenitis (with superior vena cava syndrome) SLE and AIHA CD5+ majority B lymphocytosis; reduced class-switched memory B cells; elevated DNT αβ and DNT γδ; low levels of NK cells and NK cell cytolytic activity; increased ESR, CRP, ALT, and AST; hypergammaglobulinemia; positive ANAs; anti-RNP; anti-Smith; anti-SSA Steroids (prednisone) and sirolimus 7 years Alive 
Kiykim et al., 2015, PT6 (11) Infancy Turkish c.742G>A Recurrent fever Pneumonia, gastroenteritis, and CMV Hepatosplenomegaly and lymphadenopathy SLE Naive majority B lymphocytosis and CD21low, reduced levels of switched memory B cells, increased IgM, and reduced NK functionality Ganciclovir, IVIG, topical steroids, hydroxychloroquine,
IVIG, and prophylactic antibiotics 
3.5 years Alive 
Lei et al., 2018, PT II-1, PT II-2, and PT 11-3 (12) 1 year Endogamous Pakistani c.1294G>T Constitutional symptoms (intermittent fever, night sweats, fatigue), severe thrombocytopenia, photosensitive and petechial rash, scarring alopecia, and hepatosplenomegaly NR Splenomegaly and lymphadenopathy SLE (acuta cutaneous lupus, oral/nasal ulcers) AIHA, thrombocytopenia, leukopenia, positive ANA (1:1,280), anti-dsDNA, antiphospholipid antibody, low complement, elevated CRP and ESR, elevated IgG, and low levels of C4 Steroid, rituximab (discontinued for adverse reaction),
IVIG, hydroxychloroquine,
Azathioprine, and
Ofatumumab 
NR Alive 
1 year Endogamous Pakistani c.1294G>T Constitutional symptoms (intermittent fever, night sweats, fatigue), severe thrombocytopenia, photosensitive and petechial rash, scarring alopecia, hepatosplenomegaly NR Hepatosplenomegaly, lymphadenopathy SLE (acute cutaneous lupus, serositis, and renal impairment) AIHA, thrombocytopenia, leukopenia, slight neutropenia, positive ANA (1:1,280), anti-dsDNA, antiphospholipid antibody, and low complement, elevated CRP and ESR, elevated albumin/creatinine ratio, hypergammaglobulinemia, and low levels of C4 Steroid,
IVIG, rituximab (discontinued for adverse reaction),h
ydroxychloroquine,
Azathioprine, and
Ofatumumab 
NR Alive 
26 mo Endogamous Pakistani c.1294G>T Constitutional symptoms (intermittent fever, night sweats, fatigue), severe thrombocytopenia, photosensitive and petechial rash, scarring alopecia, and hepatosplenomegaly NR Hepatosplenomegaly SLE (acute cutaneous lupus) AIHA, thrombocytopenia, positive ANA (1:1,280), anti-dsDNA, anti-Smith antibody, and low complement Steroid (IV methylprednisolone),
IVIG, and rituximab 
NR Alive 
Sharifinejad et al., 2020, PT10 (13) Infancy Afghan c.1293_1294insA Recurrent oral candidiasis, nonsevere upper respiratory and gastrointestinal tract infections, and food allergy Pneumonia, oral candidiasis, and recurrent gastroenteritis Hepatosplenomegaly and lymphadenopathy SLE Increased DNT, reduced NK number, increased IgG, and nonprotective antidiphtheria Corticosteroids, antibiotic prophylaxis 13 years Alive 
Neehus et al., 2021, PT13-17 (14) 1 year Turkish c.571+2dup BCGosis, fever, left axillary, retroauricular, and cervical lymphadenopathies BCGosis, Salmonella bacteremia, Staphylococcus aureus lymphadenitis, and recurrent gingivitis Lymphadenopathy and hepatosplenomegaly SLE Elevated naive B cells and reduced B memory, with elevated CD21low and transitional cells, hypogammaglobulinemia, ANA+, and anti-dsDNA+ Steroids
IVIG
Antimycobacterial (ceftriaxone); antibiotic prophylaxis (amoxicillin) 
14 years Alive 
7 mo Turkish c.1384C>T Fever, diarrhea, hepatosplenomegaly, failure to thrive, left axillary lymphadenopathy with purulent discharge, and BCGitis BCGitis, recurrent gastroenteritis Hepatosplenomegaly AIHA Impaired DHR test and ANA+ Steroids
Isoniazid
Rifampicin 
10 years Alive 
7 mo Turkish c.1384C>T BCGitis BCGitis, recurrent gastroenteritis and shingles Lymphadenopathy and hepatosplenomegaly AIHA Impaired DHR test and ANA+ Steroids
Isoniazid
Rifampicin
IVIG 
7 years Alive 
1 year Irani c.642del BCGitis BCGitis Hepatosplenomegaly and lymphadenopathy None Impaired DHR test and nonprotective antibody responses to tetanus and diphtheria Isoniazid
Rifampicin Azithromycin 
3 years Alive 
NR Irani c.642del Asymptomatic None None None Impaired DHR test No treatment 6 years Alive 
Gu et al., 2021, PT16-17 (15) and Jefferson et al., 2023 (3) 6 y Chinese c.36T>G Lymphadenopathy, splenomegaly, and recurrent infections NR Splenomegaly and lymphadenopathy Pancytopenia Positive antibodies for EBV, CMV, herpes simplex virus, and rubella viruses, increased TCRαβ DNT, increased CD8+ T cells and IgG, and low levels of NK cells Steroids, IRT, and sirolimus 10 years Alive 
NR Chinese c.36T>G Lymphadenopathy, splenomegaly, and anemia NR Splenomegaly and lymphadenopathy Anemia NR NR NR NR 
Yang et al., 2022, PT18 (18) 5 mo Chinese c.661C>T Recurrent fever, respiratory tract infections, and lymphadenopathy URTIs and
BCGosis 
Multiple lymphadenopathy (abdominal mesenteric lymph nodes invading the abdominal wall), bone marrow hyperplasia, and hepatosplenomegaly NR Increased DNT, elevated CD8+ T cells, low levels of CD4+ T cells, and CRP positive Isoniazid, rifampicin, ethambutol, sirolimus,
IFN-γ, and antibiotics 
8 years Alive 
Neehus et al., 2022, PT19 (19) 2 years American c.285C>A/c.376C>T URTI, lymphadenitis Burkholderia cepacia and Mycobacterium lentiflavum lymphadenopathy Infectious lymphadenopathy None Low levels of CD4+ T cells, decreased relative proportions of CD27+IgD+ and CD27+IgD memory B cells, and abnormally low DHR test Cotrimoxazole, isoniazid, and
Rifampicin 
5 years Alive 
Roderick et al., 2023, PT20-21 (9) 9 mo British c.788-2A>G/c.571C>T Listeria meningitis Listeria meningitis, Enterococcus faecium bacteremia, varicella and shingles, invasive Candida infection, recurrent episodes of Achromobacter xylosoxidans cervical lymphadenitis, and early-onset enterocolitis Lymphoid hyperplasia and splenomegaly SLE, AIHA, and ITP Increased DNTs, low B cells with low IgG with high IgM, positive anticardiolipin antibodies, Coombs positive, and positive thyroid peroxidase antibodies Steroids,
IVIG, rituximab,
Sirolimus,
Hydroxychloroquine,
HSCT, and cyclosporine (for GVHD prophylaxis) 
NR Alive 
20 mo British c.788-2A>G/c.571C>T Sepsis due to nontypeable Haemophilus influenzae Early-onset enterocolitis Lymphoid hyperplasia, splenomegaly SLE Increased DNTs, low B cells with low IgG and high IgM, positive anticardiolipin antibodies, and GAD antibody-positive IVIG replacement therapy, rituximab, sirolimus (stopped because of oral ulceration),
MMF, and hydroxychloroquine
HSCT 
8.5 years Alive 

AIHA, autoimmune hemolytic anemia; ALT, alanine aminotransferase; ANA, antinuclear antibody; AST, aspartate aminotransferase; BCG, Bacillus Calmette-Guérin; CMV, cytomegalovirus; CNS, central nervous system; CRP, C-reactive protein; C3; complement component 3; C4; complement component 4; DHR, dihydrorhodamine; DNTs, double-negative T cells; dsDNA, double-stranded DNA; EBV, Epstein-Barr virus; ESR, erythrocyte sedimentation rate; F, female; GAD, glutamic acid decarboxylase; GVHD, graft-versus-host disease; IFN-γ, interferon γ; IRT, immunoglobulin replacement therapy; ITP, immune thrombocytopenia; IVIG, intravenous immunoglobulin; LRTI, lower respiratory tract infection; M, male; MMF, mycophenolate mofetil; NK, natural killer; NR, not reported; PT, patient; RNP, ribonucleoprotein; SLE, systemic lupus erythematous; SSA, Sjögren syndrome type A; TCR, T cell receptor; URTI, upper respiratory tract infections; UTI, urinary tract infection; WHO, World Health Organization.

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