Table 1.

Summary of genetic techniques used for IEI diagnosis. CNV: copy-number variant; NGS: next-generation sequencing; SVs: structural variants; TGPs: targeted gene panels; VUS: variants of uncertain significance; WES: whole-exome sequencing; WGS: whole-genome sequencing

TechniqueFunctionKey capabilitiesLimitationsDiagnostic yieldBest use
Sanger sequencing Targeted single-gene sequencing Variant validation; known mutations; family screening for known defects One gene at a time; cannot detect large deletions/duplications Validation tool Confirming specific variants, family cascade testing, and validation of NGS findings 
TGPs Pathway-specific multigene analysis Cost-effective for specific pathways; faster than WES/WGS; good depth of coverage Limited to preselected genes; lower yield than WES/WGS 10–25% Known pathway abnormalities; initial screening 
CMA Large CNV detection Detects large CNVs (≥20–50 kb targeted, ≥100–250 kb nontargeted); identifies aneuploidies; detects microdeletions/microduplications; unbalanced rearrangements Cannot detect small CNVs (<10–20 kb); no balanced rearrangements; no point mutation detection +15% when combined with WES Syndromic presentation; complementary to WES 
WES Coding regions Analyzes ∼20,000 genes simultaneously; detects SNVs and small indels; enables gene discovery; periodic reanalysis possible; high sensitivity for coding variants No intronic/regulatory variants; limited CNV detection; poor coverage of repetitive regions; VUS interpretation challenges 25–40% (IEI cases); 30–35% (complex cases); 36–51% (critically ill neonates); up to 50% with CMA Complex/unknown IEI; negative targeted panel; phenotypically diverse presentation; gene discovery 
WGS—short-read sequencing Comprehensive genome-wide analysis Detects coding, intronic, and regulatory variants; better CNV and SV detection than WES Expensive; complex data interpretation; struggles with highly repetitive regions; large VUS burden 43%; superior to WES for undiagnosed cases WES-negative cases; regulatory variants; complex SVs 
Long-read sequencing (WGS) Ultra-long-read genome analysis Superior repeat expansion detection; resolves highly repetitive regions; better SV detection; identifies variants in previously inaccessible regions Not yet standard in clinical practice; higher cost; limited clinical validation; requires specialized expertise; lower throughput than short-read sequencing Better than short-read WGS; identifies novel variants missed by other methods Short-read negative cases; suspected expansions; complex SVs; research applications 
RNA-seq Transcriptome analysis Validates splice-site variant effects; detects aberrant splicing; identifies allelic imbalance and exon skipping; measures gene expression levels; reveals monoallelic expression Tissue-specific expression patterns; cannot detect all DNA variants; temporal expression variability Resolves ∼35% of VUS cases; functional validation tool Validating variants; explaining splicing defects; complementing DNA sequencing 
TechniqueFunctionKey capabilitiesLimitationsDiagnostic yieldBest use
Sanger sequencing Targeted single-gene sequencing Variant validation; known mutations; family screening for known defects One gene at a time; cannot detect large deletions/duplications Validation tool Confirming specific variants, family cascade testing, and validation of NGS findings 
TGPs Pathway-specific multigene analysis Cost-effective for specific pathways; faster than WES/WGS; good depth of coverage Limited to preselected genes; lower yield than WES/WGS 10–25% Known pathway abnormalities; initial screening 
CMA Large CNV detection Detects large CNVs (≥20–50 kb targeted, ≥100–250 kb nontargeted); identifies aneuploidies; detects microdeletions/microduplications; unbalanced rearrangements Cannot detect small CNVs (<10–20 kb); no balanced rearrangements; no point mutation detection +15% when combined with WES Syndromic presentation; complementary to WES 
WES Coding regions Analyzes ∼20,000 genes simultaneously; detects SNVs and small indels; enables gene discovery; periodic reanalysis possible; high sensitivity for coding variants No intronic/regulatory variants; limited CNV detection; poor coverage of repetitive regions; VUS interpretation challenges 25–40% (IEI cases); 30–35% (complex cases); 36–51% (critically ill neonates); up to 50% with CMA Complex/unknown IEI; negative targeted panel; phenotypically diverse presentation; gene discovery 
WGS—short-read sequencing Comprehensive genome-wide analysis Detects coding, intronic, and regulatory variants; better CNV and SV detection than WES Expensive; complex data interpretation; struggles with highly repetitive regions; large VUS burden 43%; superior to WES for undiagnosed cases WES-negative cases; regulatory variants; complex SVs 
Long-read sequencing (WGS) Ultra-long-read genome analysis Superior repeat expansion detection; resolves highly repetitive regions; better SV detection; identifies variants in previously inaccessible regions Not yet standard in clinical practice; higher cost; limited clinical validation; requires specialized expertise; lower throughput than short-read sequencing Better than short-read WGS; identifies novel variants missed by other methods Short-read negative cases; suspected expansions; complex SVs; research applications 
RNA-seq Transcriptome analysis Validates splice-site variant effects; detects aberrant splicing; identifies allelic imbalance and exon skipping; measures gene expression levels; reveals monoallelic expression Tissue-specific expression patterns; cannot detect all DNA variants; temporal expression variability Resolves ∼35% of VUS cases; functional validation tool Validating variants; explaining splicing defects; complementing DNA sequencing 

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