Table 1.

Clinical features, treatment responses, and outcomes in TBK1-deficient patients

IndividualFamily 1Family 2Family 3Family 4Family 5Family 6Family 7
P1P2P3P4P5P6P7P8P9P10
TBK1 variant p.W619* p.W619* p.Y212D p.R440* p. R574Sfs*11 p. R574Sfs*11 p. R574Sfs*11 p.R440* p.T77_L180del p.L717Sfs*18 
cDNA annotation c.1857G>A c.1857G>A c.634T>G c.1318C>T c.1760+4_1760+7del c.1760+4_1760+7del c.1760+4_1760+7del c.1318C>T c.228+?_540-?del c.2150_2153del 
Transcript reference (GRCh38) g.64497045 G>A g.64497045 G>A g.64474323 T>G g.64485995 C>T g. 64496403-ATAAG-A g. 64496403-ATAAG-A g. 64496403-ATAAG-A g.64485995 C>T g.64463877_64473125del g.64501340 TTAAC>T 
gnomAD (v.4.1.0) frequency 6.21E-07 6.21E-07 Private 1.89E-06 7.94E-07 7.94E-07 7.94E-07 1.89E-06 Private Private 
Predicted consequence/CADD PHRED (GRCh38-v1.7; MSC = 10.854) 39 39 30 36 Predicted LoF (frameshift) Predicted LoF (frameshift) Predicted LoF (frameshift) 36 Predicted LoF (deletion of exons 4–5) Predicted LoF (frameshift) 
Age at last evaluation (years)/Sex 34/F 26/M 7/M 11.9/F 2.4/F 1.6/F 3.5/F 4.5/M 19/M 5/F 
Systemic features 
Recurrent fever − − 
Arthritis/joint swelling + with contractures + oligoarticular, mostly distal/small joints + episodic, oligoarticular, small and large joints + polyarthritis, recurrent, self-limited 
Rash/urticarial or pustular lesions − − + erythematous + maculopapular rash − + exanthema + urticarial rash, pustulosis, neutrophilic dermatosis + urticarial rash 
Vasculitis − − + IgG-vasculitis like − − − − − 
 Lymphadenopathy/organomegaly − − − − − − − − 
Aphthous ulcers/mucosal involvement − − − − − − − 
Growth failure/poor growth − − − − − 
Other systemic (GI, renal, etc.) issues + celiac disease + abdominal pain + nephrotic syndrome − − − − − + abdominal pain 1–2×/mo, since 2016, without diarrhea and fever − 
Neurologic features 
Developmental delay/regression − + cognitive and motor retardation − − − 
Seizures (type/frequency) + convulsion (1 year of age), generalized seizure (24 years of age) + one episode of seizures with status epilepticus + febrile convulsion − + several tonic-clonic seizures + generalized seizure + seizures from 1 mo of age, initially myoclonus and fixed gaze; status epilepticus at 3 and 6 mo; later focal epilepsy with absence seizures − 
 Autoimmune/autoinflammatory) encephalitis − − − − − − − − − 
MRI/CT findings + frontal/temporal infarction involving basal ganglia due to cerebral intermediate-vessel vasculitis; hyperdense lesions in basal ganglia and corona radiata + multiple white matter lesions in brain and spinal cord Normal + cerebral and white matter atrophy with ventriculomegaly, bilateral thalamic lesions, corpus collosum thinning, and right hippocampal sclerosis; stable chronic encephalomalacic changes + brain atrophy, ventriculomegaly/hydrocephalus, lenticular/thalamic lesions, maculopathy + brain atrophy, ventriculomegaly, lenticular/thalamic lesions + mild diffuse cerebral edema, hyperdense internal veins, and bilateral thalami hypodensities − + hippocampal sclerosis; white matter abnormalities (periventricular, parietal); thalamic atrophy; cavitary sequelae of both lenticular nuclei and the right pulvinar − 
Other + stenosis/occlusion of carotid artery; frontal gliosis + MS-like disease ​ ​ + microcephaly + microcephaly, bilateral central visual impairment, delayed motor development, persistent urinary incontinence + urinary incontinence − + microcephaly (z −3); impaired fine motor skills; bilateral maculopathy − 
Viral encephalitis/infections 
Viral encephalitis − − − + ANEC and ARDS triggered by influenza A/H1N1 + influenza B + influenza B, hemorrhagic encephalitis − − − − 
Other overt viral infections + two episodes of severe rhabdomyolysis following viral infection with undefined enterovirus − − − − + CMV (5 mo) − + influenza + recurrent HSV-1 (up to 2×/mo, 2010–2018; cheek, labial, oral) − 
Vaccine tolerance/adverse reactions − − − − − − − Not reported − − 
Treatment and response 
Corticosteroids − 
Anakinra/Canakinumab − − − − Not reported − − − 
 Anti-TNF (adalimumab, infliximab, etanercept) − Not reported 
DMARDs/immuno-suppressants (MTX, AZA, THAL, COL) + MTX, AZA, THAL, COL − − − − − + MTX + MTX + COL + MTX 
Antivirals − − − + enfluvir − − + acyclovir − + acyclovir − 
Treatment efficacy Was responsive to infliximab Adverse allergic reaction to infliximab and adalimumab; systemic symptoms partially controlled with anakinra and prednisone ​ Systemic symptoms partially responsive to canakinumab; minimal corticosteroid response; encephalitic episodes refractory Unresponsive to corticosteroids, partial resolution of fever, arthritis, and skin involvement following treatment with adalimumab Good initial response to corticosteroids; partial improvement of arthritis and fever under adalimumab therapy; severe neurologic damage following influenza B–associated encephalitis Prednisolone and methotrexate resolved arthritis Systemic disease partially controlled with adalimumab; neurologically stable Partial response to adalimumab with overall improvement, but persistence of brief articular pain episodes and low-grade inflammatory syndrome; neurologic features persist Arthritis flares under etanercept and acute phase reactants remain elevated, switching to adalimumab 
Outcome 
Alive ​ ​ ​ ​ ​ 
Deceased (cause) + cause unknown ​ ​ + from influenza A–associated ADEM + from influenza B–associated encephalitis ​ + SARS-CoV-2–asscoiated multi-organ insufficiency ​ ​ ​ 
Lost to follow-up ​ ​ ​ ​ ​ ​ ​ ​ ​ 
IndividualFamily 1Family 2Family 3Family 4Family 5Family 6Family 7
P1P2P3P4P5P6P7P8P9P10
TBK1 variant p.W619* p.W619* p.Y212D p.R440* p. R574Sfs*11 p. R574Sfs*11 p. R574Sfs*11 p.R440* p.T77_L180del p.L717Sfs*18 
cDNA annotation c.1857G>A c.1857G>A c.634T>G c.1318C>T c.1760+4_1760+7del c.1760+4_1760+7del c.1760+4_1760+7del c.1318C>T c.228+?_540-?del c.2150_2153del 
Transcript reference (GRCh38) g.64497045 G>A g.64497045 G>A g.64474323 T>G g.64485995 C>T g. 64496403-ATAAG-A g. 64496403-ATAAG-A g. 64496403-ATAAG-A g.64485995 C>T g.64463877_64473125del g.64501340 TTAAC>T 
gnomAD (v.4.1.0) frequency 6.21E-07 6.21E-07 Private 1.89E-06 7.94E-07 7.94E-07 7.94E-07 1.89E-06 Private Private 
Predicted consequence/CADD PHRED (GRCh38-v1.7; MSC = 10.854) 39 39 30 36 Predicted LoF (frameshift) Predicted LoF (frameshift) Predicted LoF (frameshift) 36 Predicted LoF (deletion of exons 4–5) Predicted LoF (frameshift) 
Age at last evaluation (years)/Sex 34/F 26/M 7/M 11.9/F 2.4/F 1.6/F 3.5/F 4.5/M 19/M 5/F 
Systemic features 
Recurrent fever − − 
Arthritis/joint swelling + with contractures + oligoarticular, mostly distal/small joints + episodic, oligoarticular, small and large joints + polyarthritis, recurrent, self-limited 
Rash/urticarial or pustular lesions − − + erythematous + maculopapular rash − + exanthema + urticarial rash, pustulosis, neutrophilic dermatosis + urticarial rash 
Vasculitis − − + IgG-vasculitis like − − − − − 
 Lymphadenopathy/organomegaly − − − − − − − − 
Aphthous ulcers/mucosal involvement − − − − − − − 
Growth failure/poor growth − − − − − 
Other systemic (GI, renal, etc.) issues + celiac disease + abdominal pain + nephrotic syndrome − − − − − + abdominal pain 1–2×/mo, since 2016, without diarrhea and fever − 
Neurologic features 
Developmental delay/regression − + cognitive and motor retardation − − − 
Seizures (type/frequency) + convulsion (1 year of age), generalized seizure (24 years of age) + one episode of seizures with status epilepticus + febrile convulsion − + several tonic-clonic seizures + generalized seizure + seizures from 1 mo of age, initially myoclonus and fixed gaze; status epilepticus at 3 and 6 mo; later focal epilepsy with absence seizures − 
 Autoimmune/autoinflammatory) encephalitis − − − − − − − − − 
MRI/CT findings + frontal/temporal infarction involving basal ganglia due to cerebral intermediate-vessel vasculitis; hyperdense lesions in basal ganglia and corona radiata + multiple white matter lesions in brain and spinal cord Normal + cerebral and white matter atrophy with ventriculomegaly, bilateral thalamic lesions, corpus collosum thinning, and right hippocampal sclerosis; stable chronic encephalomalacic changes + brain atrophy, ventriculomegaly/hydrocephalus, lenticular/thalamic lesions, maculopathy + brain atrophy, ventriculomegaly, lenticular/thalamic lesions + mild diffuse cerebral edema, hyperdense internal veins, and bilateral thalami hypodensities − + hippocampal sclerosis; white matter abnormalities (periventricular, parietal); thalamic atrophy; cavitary sequelae of both lenticular nuclei and the right pulvinar − 
Other + stenosis/occlusion of carotid artery; frontal gliosis + MS-like disease ​ ​ + microcephaly + microcephaly, bilateral central visual impairment, delayed motor development, persistent urinary incontinence + urinary incontinence − + microcephaly (z −3); impaired fine motor skills; bilateral maculopathy − 
Viral encephalitis/infections 
Viral encephalitis − − − + ANEC and ARDS triggered by influenza A/H1N1 + influenza B + influenza B, hemorrhagic encephalitis − − − − 
Other overt viral infections + two episodes of severe rhabdomyolysis following viral infection with undefined enterovirus − − − − + CMV (5 mo) − + influenza + recurrent HSV-1 (up to 2×/mo, 2010–2018; cheek, labial, oral) − 
Vaccine tolerance/adverse reactions − − − − − − − Not reported − − 
Treatment and response 
Corticosteroids − 
Anakinra/Canakinumab − − − − Not reported − − − 
 Anti-TNF (adalimumab, infliximab, etanercept) − Not reported 
DMARDs/immuno-suppressants (MTX, AZA, THAL, COL) + MTX, AZA, THAL, COL − − − − − + MTX + MTX + COL + MTX 
Antivirals − − − + enfluvir − − + acyclovir − + acyclovir − 
Treatment efficacy Was responsive to infliximab Adverse allergic reaction to infliximab and adalimumab; systemic symptoms partially controlled with anakinra and prednisone ​ Systemic symptoms partially responsive to canakinumab; minimal corticosteroid response; encephalitic episodes refractory Unresponsive to corticosteroids, partial resolution of fever, arthritis, and skin involvement following treatment with adalimumab Good initial response to corticosteroids; partial improvement of arthritis and fever under adalimumab therapy; severe neurologic damage following influenza B–associated encephalitis Prednisolone and methotrexate resolved arthritis Systemic disease partially controlled with adalimumab; neurologically stable Partial response to adalimumab with overall improvement, but persistence of brief articular pain episodes and low-grade inflammatory syndrome; neurologic features persist Arthritis flares under etanercept and acute phase reactants remain elevated, switching to adalimumab 
Outcome 
Alive ​ ​ ​ ​ ​ 
Deceased (cause) + cause unknown ​ ​ + from influenza A–associated ADEM + from influenza B–associated encephalitis ​ + SARS-CoV-2–asscoiated multi-organ insufficiency ​ ​ ​ 
Lost to follow-up ​ ​ ​ ​ ​ ​ ​ ​ ​ 

LoF, loss of function; GI, gastrointestinal; MRI, magnetic resonance imaging; CT, computed tomography; ANEC, acute necrotizing encephalopathy of childhood; ARDS, acute respiratory distress syndrome; CMV, cytomegalovirus; ADEM, acute disseminated encephalomyelitis; DMARDs, disease-modifying antirheumatic drugs; MTX, methotrexate; AZA, azathioprine; THAL, thalidomide; COL, colchicine; HSV-1, herpes simplex virus type 1; CADD, combined annotation dependent depletion; PHRED, Phred-scaled CADD score; MSC, mutation significance cutoff.

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