Table 1.

Hurdles for activation of tumor-specific immunity in vivo and approaches to address thema

HurdleApproachesRecent results
Quality of therapy-induced T cells -Optimize priming signals by including costimulation and cytokines Optimal pathway to trigger and molecule use TBD 
-Evaluate NeoAg targets and “dark” antigens Single cell analysis and TCR-based tracking is allowing identification of antigen-specific T cells; broader antigen classes being tested 
T cell polyfunctionality and exhaustion -Evaluate truncal NeoAgs and more recently expressed “branch” antigen targets Single cell analysis and TCR-based tracking is allowing identification and functional profiling of antigen-specific T cells 
-Optimize priming and/or boosting signals by including costimulation and cytokines Optimal pathway and molecule use TBD, IL-12 continues to be evaluated 
Vaccine trafficking to tumor and tumor tissue penetration -Inject the tumor or tumor bed with an activating signal (chemokine, oncolytic virus, and/or stimulatory tumor killing agent) to optimize tumor targeting Oncolytic viruses continue to be developed, cocktails of costimulation with checkpoint blockade have shown distant tumor regressions 
Heterogeneity of antigen expression -Include multiple antigens in the T cell activating drugs Multiple antigens increasingly included in vaccines and NeoAg therapeutics (up to >200) 
-Promote epitope spreading More commonly evaluated in trials 
Antigen loss or MHC loss -Include multiple antigens presented by multiple human lymphocyte antigen molecules Multiple antigens increasingly included in vaccines 
-Promote epitope spreading More commonly evaluated in trials 
-Provide intratumoral IFN-γ signal to up-regulate MHC class I Optimal pathway and molecule use TBD 
HurdleApproachesRecent results
Quality of therapy-induced T cells -Optimize priming signals by including costimulation and cytokines Optimal pathway to trigger and molecule use TBD 
-Evaluate NeoAg targets and “dark” antigens Single cell analysis and TCR-based tracking is allowing identification of antigen-specific T cells; broader antigen classes being tested 
T cell polyfunctionality and exhaustion -Evaluate truncal NeoAgs and more recently expressed “branch” antigen targets Single cell analysis and TCR-based tracking is allowing identification and functional profiling of antigen-specific T cells 
-Optimize priming and/or boosting signals by including costimulation and cytokines Optimal pathway and molecule use TBD, IL-12 continues to be evaluated 
Vaccine trafficking to tumor and tumor tissue penetration -Inject the tumor or tumor bed with an activating signal (chemokine, oncolytic virus, and/or stimulatory tumor killing agent) to optimize tumor targeting Oncolytic viruses continue to be developed, cocktails of costimulation with checkpoint blockade have shown distant tumor regressions 
Heterogeneity of antigen expression -Include multiple antigens in the T cell activating drugs Multiple antigens increasingly included in vaccines and NeoAg therapeutics (up to >200) 
-Promote epitope spreading More commonly evaluated in trials 
Antigen loss or MHC loss -Include multiple antigens presented by multiple human lymphocyte antigen molecules Multiple antigens increasingly included in vaccines 
-Promote epitope spreading More commonly evaluated in trials 
-Provide intratumoral IFN-γ signal to up-regulate MHC class I Optimal pathway and molecule use TBD 

TBD, to be determined.

a

Updated from Adamik and Butterfield (2022).

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