Table 1.

TNFRSF and the corresponding ligands involved in the activation of the alternative NF-κB pathway across various human tissues and cell types

Ligand (gene)Tissue expressionCellular expressionReceptor (gene)Tissue expressionCellular expressionReference
CD40L or CD154 (CD40LGSLOs and thymus Leukocytes: activated T cells (CD4+, CD8+, and γδ), NKT, MAIT, Tregs, also detected on NK cells, mast cells, basophils, and eosinophils CD40 or TNFRSF5 (CD40SLOs and thymus Leukocytes: B cells, dendritic cells, monocytes, and macrophages.
Non-hematopoietic cells: epithelial cells (mTECs), follicular dendritic cells, endothelial cells, and fibroblasts 
(11, 12) 
LTα1β2 (LTA and LTBSLOs, thymus, and bone marrow Leukocytes: activated B and T cells, NKT, MAIT, Tregs, and dendritic cells LTβR or TNFRSF3 (LTBRSLOs, thymus, and bone marrow Leukocytes: dendritic cells, monocytes, and macrophages.
Non-hematopoietic cells: LN stromal cells (FRCs, including MRCsa), epithelial cells, endothelial cells, and mesenchymal cells 
(12, 13, 14) 
LIGHT (TNFSF14Broad, including bone marrow, SLOs, and liver Leukocytes: T cells, NK cells, macrophages, monocytes, and dendritic cells 
LIGHT-R or HVEM (TNFRSF14Broad, including SLOs B and T lymphocytes, NK cells, monocytes, macrophages, and dendritic cells (12, 15) 
RANKL (TNFSF11SLOs, bone marrow, bones, and thymus Leukocytes: activated T and B cells and NK cells.
Non-hematopoietic cells: osteoblasts, osteocytes, LN stromal cells (MRCsa), epithelial, and germ cells 
RANK (TNFRSF11AGastrointestinal tract, SLOs, and bones Hematopoietic cells: osteoclasts, macrophages, dendritic cells, and NK cells.
Non-hematopoietic cells: thymic epithelial cells (mTECs), intestinal epithelial cells (enterocytes, microfold cells), lymphatic and endothelial cells (LEC) 
(12, 14, 16, 17) 
BAFF or BLys (TNFSF13BSLOs and bone marrow Leukocytes: monocytes, macrophages, dendritic cells, and neutrophils.
Non-hematopoietic cells: LN stromal cells (FRCs, including TRCsa and MRCsa
BAFFR (TNFRSF13CSLOs and bone marrow Leukocytes: mature B cells (transitional, naïve, GC, and memory B cells) (12, 14, 18, 19) 
TWEAK (TNFSF12Broad, including bone marrow, SLOs, and liver Leukocytes: T and Ba cells, NK cells, monocytes, macrophages, dendritic cells, and neutrophils
Non-hematopoietic cells: epithelial cells and endothelial cells 
Fn14 or TWEAK-R (TNFRSF12ABroad, including SLOs Non-hematopoietic cells: epithelial cells (12, 20) 
Ligand (gene)Tissue expressionCellular expressionReceptor (gene)Tissue expressionCellular expressionReference
CD40L or CD154 (CD40LGSLOs and thymus Leukocytes: activated T cells (CD4+, CD8+, and γδ), NKT, MAIT, Tregs, also detected on NK cells, mast cells, basophils, and eosinophils CD40 or TNFRSF5 (CD40SLOs and thymus Leukocytes: B cells, dendritic cells, monocytes, and macrophages.
Non-hematopoietic cells: epithelial cells (mTECs), follicular dendritic cells, endothelial cells, and fibroblasts 
(11, 12) 
LTα1β2 (LTA and LTBSLOs, thymus, and bone marrow Leukocytes: activated B and T cells, NKT, MAIT, Tregs, and dendritic cells LTβR or TNFRSF3 (LTBRSLOs, thymus, and bone marrow Leukocytes: dendritic cells, monocytes, and macrophages.
Non-hematopoietic cells: LN stromal cells (FRCs, including MRCsa), epithelial cells, endothelial cells, and mesenchymal cells 
(12, 13, 14) 
LIGHT (TNFSF14Broad, including bone marrow, SLOs, and liver Leukocytes: T cells, NK cells, macrophages, monocytes, and dendritic cells 
LIGHT-R or HVEM (TNFRSF14Broad, including SLOs B and T lymphocytes, NK cells, monocytes, macrophages, and dendritic cells (12, 15) 
RANKL (TNFSF11SLOs, bone marrow, bones, and thymus Leukocytes: activated T and B cells and NK cells.
Non-hematopoietic cells: osteoblasts, osteocytes, LN stromal cells (MRCsa), epithelial, and germ cells 
RANK (TNFRSF11AGastrointestinal tract, SLOs, and bones Hematopoietic cells: osteoclasts, macrophages, dendritic cells, and NK cells.
Non-hematopoietic cells: thymic epithelial cells (mTECs), intestinal epithelial cells (enterocytes, microfold cells), lymphatic and endothelial cells (LEC) 
(12, 14, 16, 17) 
BAFF or BLys (TNFSF13BSLOs and bone marrow Leukocytes: monocytes, macrophages, dendritic cells, and neutrophils.
Non-hematopoietic cells: LN stromal cells (FRCs, including TRCsa and MRCsa
BAFFR (TNFRSF13CSLOs and bone marrow Leukocytes: mature B cells (transitional, naïve, GC, and memory B cells) (12, 14, 18, 19) 
TWEAK (TNFSF12Broad, including bone marrow, SLOs, and liver Leukocytes: T and Ba cells, NK cells, monocytes, macrophages, dendritic cells, and neutrophils
Non-hematopoietic cells: epithelial cells and endothelial cells 
Fn14 or TWEAK-R (TNFRSF12ABroad, including SLOs Non-hematopoietic cells: epithelial cells (12, 20) 

Bold typeface is used to indicate the principal cell type in which expression occurs. FRC, fibroblastic reticular cells; LEC, lymphatic endothelial cells; LN, lymph node; MRC, marginal reticular cells; TRC, T-zone reticular cells.

a

Evidence only at the mRNA level.

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