Table 1.

Comparison of key attributes of chronically stimulated CD4+ vs. CD8+ T cells

AttributeChronically stimulated CD8+ T cellsChronically stimulated CD4+ T cellsReferences
IR expression ↑ PD-1, CTLA4, TIM-3 and LAG-3. ↑2B4, LILRB4 ↑ PD-1, CTLA4, TIM-3 and LAG-3 Dow et al. (2013), Goding et al. (2013), Crawford et al. (2014), Pour et al. (2019), Tracy et al. (2022)  
↑ CD200 and BTLA4 
Transcriptional regulation TOX, NFAT, PRDM1, BACH2, BATF, and EOMES TOX, NFAT, PRDM1, BACH2, BATF, and EOMES Crawford et al. (2014), Trefzer et al. (2021), Sumida et al. (2022)  
Uniquely upregulates IKZF2 (Helios) 
Functional profile ↓ Self-renew ↓ Self-renew Beltra et al. (2020), Fu et al. (2020)  
↓ IFNγ, TNFα, and IL2 ↓ TNFα and IL2 but retains IFNγ, ↑ Tfh attributes (IL21 and IL10) Crawford et al. (2014), Catakovic et al. (2017), Balança et al. (2021)  
Cytotoxicity attributes ++ to +++ 0 to ++ Oh et al. (2020), Liu et al. (2022), Sultan et al. (2024)  
Epigenetic traits Characteristic epigenetic remodeling in IR, cytokine, and cytotoxicity genes Pauken et al. (2016), Schietinger et al. (2016), Stephen et al. (2017), Satpathy et al. (2019)  
Ontogeny Terminally exhausted arise from a self-renewing TCF7+ Tpex subset Arise from a self-renewing TCF7+ progenitor Beltra et al. (2020), Xia et al. (2022), Zhou et al. (2024)  
AttributeChronically stimulated CD8+ T cellsChronically stimulated CD4+ T cellsReferences
IR expression ↑ PD-1, CTLA4, TIM-3 and LAG-3. ↑2B4, LILRB4 ↑ PD-1, CTLA4, TIM-3 and LAG-3 Dow et al. (2013), Goding et al. (2013), Crawford et al. (2014), Pour et al. (2019), Tracy et al. (2022)  
↑ CD200 and BTLA4 
Transcriptional regulation TOX, NFAT, PRDM1, BACH2, BATF, and EOMES TOX, NFAT, PRDM1, BACH2, BATF, and EOMES Crawford et al. (2014), Trefzer et al. (2021), Sumida et al. (2022)  
Uniquely upregulates IKZF2 (Helios) 
Functional profile ↓ Self-renew ↓ Self-renew Beltra et al. (2020), Fu et al. (2020)  
↓ IFNγ, TNFα, and IL2 ↓ TNFα and IL2 but retains IFNγ, ↑ Tfh attributes (IL21 and IL10) Crawford et al. (2014), Catakovic et al. (2017), Balança et al. (2021)  
Cytotoxicity attributes ++ to +++ 0 to ++ Oh et al. (2020), Liu et al. (2022), Sultan et al. (2024)  
Epigenetic traits Characteristic epigenetic remodeling in IR, cytokine, and cytotoxicity genes Pauken et al. (2016), Schietinger et al. (2016), Stephen et al. (2017), Satpathy et al. (2019)  
Ontogeny Terminally exhausted arise from a self-renewing TCF7+ Tpex subset Arise from a self-renewing TCF7+ progenitor Beltra et al. (2020), Xia et al. (2022), Zhou et al. (2024)  

BATF, basic leucine zipper ATF-like transcription factor; EOMES, eomesodermin; BACH2, BTB domain and CNC homolog 2.

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