Table 1.
History table of OHC eM frequency measurements
PublicationMethod to evoke and measure load-free eMFrequency response cut-offVoltage delivery and/or eM measurement limitationsNotes
(Kachar et al., 1986) Transcellular AC current; standard video No estimate of roll-off; 1–30 Hz frame rates. Rm * Cm time constant; limited to 60 frames/s  
(Ashmore, 1987) Whole-cell voltage clamp; photodiode Dual Lorentzian cut-offs at 67 and 664 Hz Voltage clamp Rs * Cm time constant  
(Santos-Sacchi, 1992) Whole-cell voltage clamp; photodiode Single Lorentzian cut-off at 1 kHz Voltage clamp Rs * Cm time constant  
(Reuter et al., 1992) Transcellular AC current stimulation in organ explants; stroboscopic video 40 dB down at 15 kHz Membrane Rm * Cm time constant  
(Dallos and Evans, 1995) Microchamber cell partitioning, AC voltage from low-impedance waveform generator; photodiode No estimate of roll-off, as shape of frequency response is a function of apical to basal impedances; eM detection above 10 kHz; zero microchamber V offset Rs * Cin (input capacitance of partitioned cell) for command voltage; ratio of partitioned impedance modifies roll-off shape Low frequency eM gain:∼5 nm/mV indicative of offset away from Vh; no estimation of V delivery roll-off based on generated currents 
(Gale and Ashmore 1997b) On-cell and excised membrane patch under voltage clamp Patch movements (≈ eM) measured with step voltages; 10 kHz cut-off for patch NLC; 0.19× lower cut-off for movements Near ideal voltage clamp First indication that eM and NLC frequency response may differ 
(Frank et al., 1999) Microchamber cell partitioning; voltage clamp; laser Doppler vibrometer Variable cut-off; for 30 µm cell extrusion 33 kHz; smaller extrusions > 70 kHz; zero microchamber V offset Rs * Cin for command voltage; ratio of partitioned impedance modifies roll-off shape Low frequency eM gain:∼5 nm/mV indicative of offset away from Vh; voltage corrections based on Rs * (stray || membrane capacitance) 
(Kitani et al., 2011) Transcellular AC current; video analysis at 18 kHz Responses measured up to 4 kHz Rm * Cm time constant Time dependent changes in eM magnitude may be attributed to mechanisms other than prestin 
(Santos-Sacchi and Tan, 2018) Microchamber cell partitioning; voltage clamp; video analysis at 50 KHz frame rate At microchamber V offset to Vh, dual Lorentzian cut-offs of 33 Hz and 6.3 kHz; at zero microchamber V offset, dual Lorentzian cut-offs of 234 Hz and 8.7 kHz Rs * Cin for command voltage; ratio of partitioned impedance modifies roll-off shape Low frequency eM gain (at Vh): 16.8 nm/mV; voltage corrections based on exponential current decays during V stimulation at two microchamber offsets; eM measures made under whole cell voltage clamp were not corrected for delivery roll-off 
PublicationMethod to evoke and measure load-free eMFrequency response cut-offVoltage delivery and/or eM measurement limitationsNotes
(Kachar et al., 1986) Transcellular AC current; standard video No estimate of roll-off; 1–30 Hz frame rates. Rm * Cm time constant; limited to 60 frames/s  
(Ashmore, 1987) Whole-cell voltage clamp; photodiode Dual Lorentzian cut-offs at 67 and 664 Hz Voltage clamp Rs * Cm time constant  
(Santos-Sacchi, 1992) Whole-cell voltage clamp; photodiode Single Lorentzian cut-off at 1 kHz Voltage clamp Rs * Cm time constant  
(Reuter et al., 1992) Transcellular AC current stimulation in organ explants; stroboscopic video 40 dB down at 15 kHz Membrane Rm * Cm time constant  
(Dallos and Evans, 1995) Microchamber cell partitioning, AC voltage from low-impedance waveform generator; photodiode No estimate of roll-off, as shape of frequency response is a function of apical to basal impedances; eM detection above 10 kHz; zero microchamber V offset Rs * Cin (input capacitance of partitioned cell) for command voltage; ratio of partitioned impedance modifies roll-off shape Low frequency eM gain:∼5 nm/mV indicative of offset away from Vh; no estimation of V delivery roll-off based on generated currents 
(Gale and Ashmore 1997b) On-cell and excised membrane patch under voltage clamp Patch movements (≈ eM) measured with step voltages; 10 kHz cut-off for patch NLC; 0.19× lower cut-off for movements Near ideal voltage clamp First indication that eM and NLC frequency response may differ 
(Frank et al., 1999) Microchamber cell partitioning; voltage clamp; laser Doppler vibrometer Variable cut-off; for 30 µm cell extrusion 33 kHz; smaller extrusions > 70 kHz; zero microchamber V offset Rs * Cin for command voltage; ratio of partitioned impedance modifies roll-off shape Low frequency eM gain:∼5 nm/mV indicative of offset away from Vh; voltage corrections based on Rs * (stray || membrane capacitance) 
(Kitani et al., 2011) Transcellular AC current; video analysis at 18 kHz Responses measured up to 4 kHz Rm * Cm time constant Time dependent changes in eM magnitude may be attributed to mechanisms other than prestin 
(Santos-Sacchi and Tan, 2018) Microchamber cell partitioning; voltage clamp; video analysis at 50 KHz frame rate At microchamber V offset to Vh, dual Lorentzian cut-offs of 33 Hz and 6.3 kHz; at zero microchamber V offset, dual Lorentzian cut-offs of 234 Hz and 8.7 kHz Rs * Cin for command voltage; ratio of partitioned impedance modifies roll-off shape Low frequency eM gain (at Vh): 16.8 nm/mV; voltage corrections based on exponential current decays during V stimulation at two microchamber offsets; eM measures made under whole cell voltage clamp were not corrected for delivery roll-off 

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