Table II.

Higher escape rates are observed in the first T cells of HIV-1 infection

Patient Reference Mutation/sa Average rate escape, kb t50c CD8 T cell epitoped Confirmed T cell response (HXB2 position)e 
   day−1 days   
CH40 a. Gag A120D 0.17 31 no no 
CH40 b. Gag I401L, R403K 0.17 31 yes yes (Gag389-406) 
CH40 d. Pol K76R 0.015 119 yes nof 
CH40 h. Vif T167I 0.37 38 yes no 
CH40 i. Vpr S84R 0.16 31 yes no 
CH40 t. Nef S188R/N, R196Q 0.22 15 yes yes, (Nef185-202) 
CH77 l. Tat I64TS 0.42 yes nog 
CH77 s. Env R355K/S/N,T358A 0.36 yes yes, (Env350-368) 
CH77 aa. Nef R21K/G, R22K/G/,TD23E/N, P25S 0.30 19 yes yes, (Nef17-34) 
CH77 cc. Nef K82Q/T/P, A83G, L85H, L87I 0.29 24 yes yes, (Nef81-98) 
CH77 b. Gag I147L 0.063 101 yes yes, (Gag140-157) IW9 
CH77 c. Gag T242N,V247I, G248E/S 0.035 124 yes yes, (Gag236-253) TW10 
CH58 e. Env Y586H 0.10 21 yes yes, (Env576-596) 
CH58 g. Env F829V/S/L, I830M/L/F, A832T, V833I 0.12 27 no no 
CH58 b. Gag I147L 0.007 430 yes yes, (Gag140-157) IW9 
CH58 c. Gag T242N, G248E 0.085 60 yes yes, (Gag236-253), TW10 
SUMA0874  Rev R48K, Q51H, Q53R, S54L, L55I 0.32 30 yes yes, (Rev47-55) 
Median  all demonstrated T cell epitopes 0.17 (max. 0.42) 0.14 (max. 0.36)   
Patient Reference Mutation/sa Average rate escape, kb t50c CD8 T cell epitoped Confirmed T cell response (HXB2 position)e 
   day−1 days   
CH40 a. Gag A120D 0.17 31 no no 
CH40 b. Gag I401L, R403K 0.17 31 yes yes (Gag389-406) 
CH40 d. Pol K76R 0.015 119 yes nof 
CH40 h. Vif T167I 0.37 38 yes no 
CH40 i. Vpr S84R 0.16 31 yes no 
CH40 t. Nef S188R/N, R196Q 0.22 15 yes yes, (Nef185-202) 
CH77 l. Tat I64TS 0.42 yes nog 
CH77 s. Env R355K/S/N,T358A 0.36 yes yes, (Env350-368) 
CH77 aa. Nef R21K/G, R22K/G/,TD23E/N, P25S 0.30 19 yes yes, (Nef17-34) 
CH77 cc. Nef K82Q/T/P, A83G, L85H, L87I 0.29 24 yes yes, (Nef81-98) 
CH77 b. Gag I147L 0.063 101 yes yes, (Gag140-157) IW9 
CH77 c. Gag T242N,V247I, G248E/S 0.035 124 yes yes, (Gag236-253) TW10 
CH58 e. Env Y586H 0.10 21 yes yes, (Env576-596) 
CH58 g. Env F829V/S/L, I830M/L/F, A832T, V833I 0.12 27 no no 
CH58 b. Gag I147L 0.007 430 yes yes, (Gag140-157) IW9 
CH58 c. Gag T242N, G248E 0.085 60 yes yes, (Gag236-253), TW10 
SUMA0874  Rev R48K, Q51H, Q53R, S54L, L55I 0.32 30 yes yes, (Rev47-55) 
Median  all demonstrated T cell epitopes 0.17 (max. 0.42) 0.14 (max. 0.36)   

Summary of characteristics of several mutations that became fixed in the virus population within several weeks after the onset of symptoms.

a

Note, that in the majority of cases, several different nonsynonymous mutations occurred in a given peptide in the model. All such escape variants are assumed not to be recognized by the corresponding CTL responses. Analysis of the extension of this model assumption will be presented elsewhere.

b

The rate of escape, k, determines the mean rate at which mutations accumulated in the viral population.

c

t50 is the predicted time of the mutant to reach 50% frequency in the viral population.

d

The mutations occur in known CTL epitopes or in sites with CTL anchor binding motifs given the patient's HLA type (ELF software, LANL).

e

A CD8 T cell response (as measured by ELISpot or ICS) to a peptide corresponding to the transmitted virus was detected at that site.

f

This mutation is proximal to the Pol 80–97 epitope and may have had an impact on its processing. See also Fig. 5 H.

g

This region partially overlaps with Rev in another reading frame. See paragraph three of the CH77 section of the Results for further discussion.

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