Table I.

Fluorescence imaging artifacts accompanying pseudopodia formation

Cellular condition Consequence Fluorescence Imaging Artifact 
A. Non-uniformity of plasma membrane topography   
(i) Membrane ruffling   
(ii) Increased sub-resolution convolutions in pseudopodial membrane Locally increased membrane within the imaging plane Higher local fluorescence of membrane probes 
(iii) Reduced sub-resolution convolutions in pseudopodial membrane Locally reduced membrane within the imaging plane Lower local fluorescence of membrane probes 
B. Non-uniformity of pseudopodial z-plane thickness Locally reduced cytoplasmic volume within the imaging plane Lower fluorescence in “thinner” region of pseudopodium 
(i) Extended pseudopodia thinner than non-pseudopodial regions 
(ii) Thickening of termini relative of extending pseudopodia Locally increased cytoplasmic volume within the imaging plane Higher fluorescence localized to “thicker” regions of pseudopodium 
(iii) z-plane extensions at pseudopodia termini not yet attached to substrate 
C. Organelle exclusion from pseudopodiaa   
(i) Reduced light scattering in extending pseudopodia (see Fig. 2 b) (i) Locally increased excitation efficiency  
(ii) Reduced cytosolic density (see Fig. 2 a) (ii) Locally increased cytoplasmic fluor concentration Higher fluorescence in pseudopodia 
Cellular condition Consequence Fluorescence Imaging Artifact 
A. Non-uniformity of plasma membrane topography   
(i) Membrane ruffling   
(ii) Increased sub-resolution convolutions in pseudopodial membrane Locally increased membrane within the imaging plane Higher local fluorescence of membrane probes 
(iii) Reduced sub-resolution convolutions in pseudopodial membrane Locally reduced membrane within the imaging plane Lower local fluorescence of membrane probes 
B. Non-uniformity of pseudopodial z-plane thickness Locally reduced cytoplasmic volume within the imaging plane Lower fluorescence in “thinner” region of pseudopodium 
(i) Extended pseudopodia thinner than non-pseudopodial regions 
(ii) Thickening of termini relative of extending pseudopodia Locally increased cytoplasmic volume within the imaging plane Higher fluorescence localized to “thicker” regions of pseudopodium 
(iii) z-plane extensions at pseudopodia termini not yet attached to substrate 
C. Organelle exclusion from pseudopodiaa   
(i) Reduced light scattering in extending pseudopodia (see Fig. 2 b) (i) Locally increased excitation efficiency  
(ii) Reduced cytosolic density (see Fig. 2 a) (ii) Locally increased cytoplasmic fluor concentration Higher fluorescence in pseudopodia 
a

Artifacts listed in C have not previously been recognized and is the main subject of this paper.

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