Table 1.

Isometric twitch-tension parameters at basal conditions (30°C, 2 mM [Ca2+]out, 1-Hz stimulation rate) and in the presence of 10−6 M isoproterenol or high extracellular calcium (8 mM [Ca2+]out).

Parameter Atrial trabeculae Ventricular trabeculae 
Basal conditions WT (n = 12) R4496C (n = 10) P value WT (n = 12) R4496C (n = 11) P value 
Peak tension (mN/mm211.3 ± 1.3 12.9 ± 2.4 NS 26.0 ± 4.4 26.8 ± 4.5 NS 
Time to peak (ms) 44.9 ± 2.4 44.7 ± 1.7 NS 63.4 ± 4.2 64.9 ± 3.7 NS 
Time to 50% relaxation (ms) 24.1 ± 1.1 24.8 ± 2.2 NS 42.3 ± 1.7 46.1 ± 1.8 NS 
Isoproterenol       
Peak tension (mN/mm249.3 ± 8.3 22.9 ± 6.4 <0.01 66.3 ± 12.4 41.8 ± 8.5 <0.05 
Time to peak (ms) 39.3 ± 4.4 37.7 ± 3.7 NS 43.7 ± 3.6 44.9 ± 3.7 NS 
Time to 50% relaxation (ms) 17.5 ± 2.1 18.1 ± 2.2 NS 22.3 ± 1.9 23.1 ± 1.8 NS 
High extracellular calcium       
Peak tension (mN/mm247.2 ± 8.6 25.2 ± 5.6 <0.05 65.4 ± 12.2 49.1 ± 9.0 <0.05 
Time to peak (ms) 47.6 ± 3.3 48.4 ± 2.6 NS 64.1 ± 3.6 64.5 ± 4.1 NS 
Time to 50% relaxation (ms) 25.8 ± 2.7 27.1 ± 3.1 NS 45.3 ± 4.8 46.5 ± 2.3 NS 
Parameter Atrial trabeculae Ventricular trabeculae 
Basal conditions WT (n = 12) R4496C (n = 10) P value WT (n = 12) R4496C (n = 11) P value 
Peak tension (mN/mm211.3 ± 1.3 12.9 ± 2.4 NS 26.0 ± 4.4 26.8 ± 4.5 NS 
Time to peak (ms) 44.9 ± 2.4 44.7 ± 1.7 NS 63.4 ± 4.2 64.9 ± 3.7 NS 
Time to 50% relaxation (ms) 24.1 ± 1.1 24.8 ± 2.2 NS 42.3 ± 1.7 46.1 ± 1.8 NS 
Isoproterenol       
Peak tension (mN/mm249.3 ± 8.3 22.9 ± 6.4 <0.01 66.3 ± 12.4 41.8 ± 8.5 <0.05 
Time to peak (ms) 39.3 ± 4.4 37.7 ± 3.7 NS 43.7 ± 3.6 44.9 ± 3.7 NS 
Time to 50% relaxation (ms) 17.5 ± 2.1 18.1 ± 2.2 NS 22.3 ± 1.9 23.1 ± 1.8 NS 
High extracellular calcium       
Peak tension (mN/mm247.2 ± 8.6 25.2 ± 5.6 <0.05 65.4 ± 12.2 49.1 ± 9.0 <0.05 
Time to peak (ms) 47.6 ± 3.3 48.4 ± 2.6 NS 64.1 ± 3.6 64.5 ± 4.1 NS 
Time to 50% relaxation (ms) 25.8 ± 2.7 27.1 ± 3.1 NS 45.3 ± 4.8 46.5 ± 2.3 NS 

Values are expressed as means ± SEM. The p-value is estimated by the Student’s t test. The number in parentheses is the number of preparations. Time to peak is the time from stimulus to peak contraction. Time to 50% relaxation is the time from peak to 50% decay of force.

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