Table 1.

Summary of the data on the contribution of different mechanisms to cAMP compartmentalization

Cell type AC (cAMP synthesis) PDE (cAMP degradation) Hindered diffusion Cell shape cAMP buffers Notes References 
HEK293 cells ND Yes: PDE inhibition abolishes differences cytosol/membrane Yes ND Yes Modified CNG channel Rich et al. (2000, 2001, 2007) 
HEK293 cells ND Yes: Loss of gradient with PDE4 RNAi; no effect of PDE4 inhibitors No: Computational analysis ND ND PKA/EPAC reporters Terrin et al. (2006); Oliveira et al. (2010)  
HEK293 cells Yes (targeted AC8) Yes: Loss of membrane compartment after PDE4 inhibition ND ND ND Gravin-mediated complex Willoughby et al. (2006)  
MEFs ND No: PDE4B ablation increases cytosol/membrane gradient ND ND ND CNGC/EPAC reporters Blackman et al. (2011)  
Neonatal cardiac myocytes ND Yes: IBMX abolishes local cAMP accumulation ND ND ND PKA reporter Zaccolo and Pozzan (2002); Mongillo et al. (2004)  
Neonatal cardiac myocytes Yes (receptor-dependent βAR vs. PGE2) Yes: PDE inhibition decreases the PKA-phosphorylation gradients Yes ND Yes Membrane/cytosol restricted diff. local cAMP uncaging Saucerman et al. (2006)  
Neurons (dendrites)  Yes: Role of negative regulatory loop No Yes: Dendrite diameter   Bacskai et al. (1993); Neves et al. (2008)  
Adult cardiac myocytes Differences β1/β2 responses No: No PDE effect on diffusion of the β2AR restricted signal Yes ND ND HCN2-cAMP probe in transgenic mice Nikolaev et al. (2006)  
Vascular endothelium Yes: Membrane vs. cytosol AC Yes: PDE4D4 displacement affects membrane cAMP; Rolipram disrupts the gradient Yes Yes Yes Manipulation of site of cAMP synthesis Sayner et al. (2006); Feinstein et al. (2012)  
Cell type AC (cAMP synthesis) PDE (cAMP degradation) Hindered diffusion Cell shape cAMP buffers Notes References 
HEK293 cells ND Yes: PDE inhibition abolishes differences cytosol/membrane Yes ND Yes Modified CNG channel Rich et al. (2000, 2001, 2007) 
HEK293 cells ND Yes: Loss of gradient with PDE4 RNAi; no effect of PDE4 inhibitors No: Computational analysis ND ND PKA/EPAC reporters Terrin et al. (2006); Oliveira et al. (2010)  
HEK293 cells Yes (targeted AC8) Yes: Loss of membrane compartment after PDE4 inhibition ND ND ND Gravin-mediated complex Willoughby et al. (2006)  
MEFs ND No: PDE4B ablation increases cytosol/membrane gradient ND ND ND CNGC/EPAC reporters Blackman et al. (2011)  
Neonatal cardiac myocytes ND Yes: IBMX abolishes local cAMP accumulation ND ND ND PKA reporter Zaccolo and Pozzan (2002); Mongillo et al. (2004)  
Neonatal cardiac myocytes Yes (receptor-dependent βAR vs. PGE2) Yes: PDE inhibition decreases the PKA-phosphorylation gradients Yes ND Yes Membrane/cytosol restricted diff. local cAMP uncaging Saucerman et al. (2006)  
Neurons (dendrites)  Yes: Role of negative regulatory loop No Yes: Dendrite diameter   Bacskai et al. (1993); Neves et al. (2008)  
Adult cardiac myocytes Differences β1/β2 responses No: No PDE effect on diffusion of the β2AR restricted signal Yes ND ND HCN2-cAMP probe in transgenic mice Nikolaev et al. (2006)  
Vascular endothelium Yes: Membrane vs. cytosol AC Yes: PDE4D4 displacement affects membrane cAMP; Rolipram disrupts the gradient Yes Yes Yes Manipulation of site of cAMP synthesis Sayner et al. (2006); Feinstein et al. (2012)  

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