Table 1.
Clinical data of patients with ATP6AP2 mutations
ParameterPatient 1Patient 2Patient 3
Sex Male Male Male 
Current age 17 yr 21 yr 10 mo 
Consanguinity None None None 
Ethnicity Caucasian Caucasian Caucasian 
Mutation cDNA c.293C>T c.212G>A c.212G>A 
Mutation protein p.Leu98Ser (L98S) p. Arg71His (R71H) p. Arg71His (R71H) 
Glycosylation defect IEF: tetrasialotransferrin: 34.2% (norm: 48.5–65.3%); trisialotransferrin: 28.9% (norm: 5.5–15.1%); disialotransferrin: 15% (norm: 2.0–6.1%); monosialotransferrin: 3.3% (norm: 0.0–3.7%); asialotransferrin: 1.5% (norm: 0%) HPLC: tetrasialotransferrin 55.17% (norm: 85.7–94.0%); trisialotransferrin 33.11% (norm: 1.16–6.36%); disialotransferrin 8.92% (norm: 0.38–1.82%); monosialotransferrin 1.68% (norm: 0%) HPLC: tetrasialotransferrin 35.7% (norm: 85.7–94.0%); trisialotransferrin 36.4% (norm: 1.16–6.36%); disialotransferrin 18.7% (norm: 0.38–1.82%); monosialotransferrin 9.0% (norm: 0%) 
Onset of symptoms 5 mo Directly after birth Directly after birth 
Liver Prolonged neonatal jaundice and persistent hepatosplenomegaly with recurrent episodes of hypoalbuminemia and ascites No hepatomegaly in childhood, but liver parenchyma of an inhomogeneous, hyperechoic structure. Prolonged neonatal jaundice and hepatosplenomegaly. Cholestasis. Irregular liver parenchyma with portosystemic shunts and ascites. Hypoalbuminemia, hyperammonemia. Coagulopathy. Listed for liver transplantation. 
  Today: hepatomegaly, mild elevation of liver enzymes  
Liver biopsy Micronodular hepatic cirrhosis with moderate macrovesicular steatosis (biopsy at 8 mo), elevated copper Not available Diffuse micronodular hepatic cirrhosis 
   Elevated copper 
Neonatal icterus 
Splenomegaly 
Infections Recurrent severe infections (e.g., sepsis, peritonitis) Recurring pulmonary and upper respiratory tract infections throughout infancy and childhood Recurring upper respiratory tract infections, positive blood cultures with pneumococcus at one occasion 
Neurological symptoms Mild cognitive impairment Ataxic gait, mild cognitive impairment Normal 
Cutis laxa Mild to moderate Pronounced, improvement over time Pronounced 
Hypogammaglo-bulinemia 
IgG 2.31 g/L (reference: 5.6–13.8 g/L) Reduced IgG1 and IgG3 
IgM <0.17 g/L 18 mg/dl (reference: 40–230 mg/dl) 
IgA <0.08 g/L <15 mg/dl (reference: 70–400 mg/dl) 
Increased transaminases At 17 yr, mild elevation of transaminases (AST 61 U/l [reference: 15–46 UI/l], ALT 51 U/l [reference: 10–40 U/l]) Ranging from mild elevation at age 9 yr: AST 92 U/l (reference: 8–60 U/l), ALT 59 U/l (reference: <44 U/l) to pronounced hepatic affection (AST 135 U/l) Within the first year: AST 100–160 U/l, (reference: <71 U/l) 
Serum copper Not available At 1 yr: 1.5 µmol/l (reference: 10–30 µmol/l) Normal serum copper at the age of 4 mo: 89 µg/dl (reference: 65–165 µg/dl) 
  Normalized over the following months  
Serum lipids At age 17 yr (liver normal): At age 20 yr: At age 5 mo: 
 Cholesterol: 172 (reference: 200 mg/dl) Cholesterol: 199 (reference: 200 mg/dl) Cholesterol: 254 (reference: 81–147 mg/dl) 
 LDL-cholesterol: 90 (reference: 50–130 md/dl) LDL-cholesterol: 149 (reference: 50–130 md/dl)  
Other laboratory findings Low factor V and VI Low factor XI (58% [reference: >70%]) and free protein S (57.5% [reference: 60–140%]) Low factors II (27.7% [reference: 60–120%]), V (35.9% [reference: 55–130%]), VII (21.9% [reference: 47–130%]), IX (17.27% [reference: 36–136%]), and XI (23.9% [reference: 49–134%]) 
 Factor V: 30% (reference: 65–148%), factor VII: 24% (reference: 58–115%)   
Other clinical findings Mild dysmorphic features Low-set ears, micrognathia, a flat and wide-set chest, laterally facing nipples, and hypospadia  
ParameterPatient 1Patient 2Patient 3
Sex Male Male Male 
Current age 17 yr 21 yr 10 mo 
Consanguinity None None None 
Ethnicity Caucasian Caucasian Caucasian 
Mutation cDNA c.293C>T c.212G>A c.212G>A 
Mutation protein p.Leu98Ser (L98S) p. Arg71His (R71H) p. Arg71His (R71H) 
Glycosylation defect IEF: tetrasialotransferrin: 34.2% (norm: 48.5–65.3%); trisialotransferrin: 28.9% (norm: 5.5–15.1%); disialotransferrin: 15% (norm: 2.0–6.1%); monosialotransferrin: 3.3% (norm: 0.0–3.7%); asialotransferrin: 1.5% (norm: 0%) HPLC: tetrasialotransferrin 55.17% (norm: 85.7–94.0%); trisialotransferrin 33.11% (norm: 1.16–6.36%); disialotransferrin 8.92% (norm: 0.38–1.82%); monosialotransferrin 1.68% (norm: 0%) HPLC: tetrasialotransferrin 35.7% (norm: 85.7–94.0%); trisialotransferrin 36.4% (norm: 1.16–6.36%); disialotransferrin 18.7% (norm: 0.38–1.82%); monosialotransferrin 9.0% (norm: 0%) 
Onset of symptoms 5 mo Directly after birth Directly after birth 
Liver Prolonged neonatal jaundice and persistent hepatosplenomegaly with recurrent episodes of hypoalbuminemia and ascites No hepatomegaly in childhood, but liver parenchyma of an inhomogeneous, hyperechoic structure. Prolonged neonatal jaundice and hepatosplenomegaly. Cholestasis. Irregular liver parenchyma with portosystemic shunts and ascites. Hypoalbuminemia, hyperammonemia. Coagulopathy. Listed for liver transplantation. 
  Today: hepatomegaly, mild elevation of liver enzymes  
Liver biopsy Micronodular hepatic cirrhosis with moderate macrovesicular steatosis (biopsy at 8 mo), elevated copper Not available Diffuse micronodular hepatic cirrhosis 
   Elevated copper 
Neonatal icterus 
Splenomegaly 
Infections Recurrent severe infections (e.g., sepsis, peritonitis) Recurring pulmonary and upper respiratory tract infections throughout infancy and childhood Recurring upper respiratory tract infections, positive blood cultures with pneumococcus at one occasion 
Neurological symptoms Mild cognitive impairment Ataxic gait, mild cognitive impairment Normal 
Cutis laxa Mild to moderate Pronounced, improvement over time Pronounced 
Hypogammaglo-bulinemia 
IgG 2.31 g/L (reference: 5.6–13.8 g/L) Reduced IgG1 and IgG3 
IgM <0.17 g/L 18 mg/dl (reference: 40–230 mg/dl) 
IgA <0.08 g/L <15 mg/dl (reference: 70–400 mg/dl) 
Increased transaminases At 17 yr, mild elevation of transaminases (AST 61 U/l [reference: 15–46 UI/l], ALT 51 U/l [reference: 10–40 U/l]) Ranging from mild elevation at age 9 yr: AST 92 U/l (reference: 8–60 U/l), ALT 59 U/l (reference: <44 U/l) to pronounced hepatic affection (AST 135 U/l) Within the first year: AST 100–160 U/l, (reference: <71 U/l) 
Serum copper Not available At 1 yr: 1.5 µmol/l (reference: 10–30 µmol/l) Normal serum copper at the age of 4 mo: 89 µg/dl (reference: 65–165 µg/dl) 
  Normalized over the following months  
Serum lipids At age 17 yr (liver normal): At age 20 yr: At age 5 mo: 
 Cholesterol: 172 (reference: 200 mg/dl) Cholesterol: 199 (reference: 200 mg/dl) Cholesterol: 254 (reference: 81–147 mg/dl) 
 LDL-cholesterol: 90 (reference: 50–130 md/dl) LDL-cholesterol: 149 (reference: 50–130 md/dl)  
Other laboratory findings Low factor V and VI Low factor XI (58% [reference: >70%]) and free protein S (57.5% [reference: 60–140%]) Low factors II (27.7% [reference: 60–120%]), V (35.9% [reference: 55–130%]), VII (21.9% [reference: 47–130%]), IX (17.27% [reference: 36–136%]), and XI (23.9% [reference: 49–134%]) 
 Factor V: 30% (reference: 65–148%), factor VII: 24% (reference: 58–115%)   
Other clinical findings Mild dysmorphic features Low-set ears, micrognathia, a flat and wide-set chest, laterally facing nipples, and hypospadia  

IEF, isoelectric focusing.

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