Table 1.

Genomic abnormalities observed in mouse and human PSCs

Aberration type Mouse PSCs Human PSCs Comparison of human and mouse PSCs Origin Gene enrichment Likely mechanism of formation 
Chromosomal aberrations Recurrent aberrations Gains 8 and 11Deletions 10qB and 14q(a, b, c, d, e, f) Gains 1, 12, 17, 20 and X (g, h, i, j, k, l, m, n) Human chromosome 17 is completely syntenic to the distal half of mouse chromosome 11(c) Most aberrations arise in culture during propagation (culture adaptation) (i, n) Difficult to analyze, as aberrations contain multiple genes Defects in chromosomal segregation during cell division 
Frequency of aberrations ∼38% in mESCs ∼23% in miPSCs (c) ∼25% of the aberrations involve chromosomes 8 and 11 (c) ∼32–34% in hESCs (i, n) ∼20% in hiPSCs (n) ∼50% of the aberrations involve chromosomes 1,12,17 or 20 (i) 
Subchromosomal aberrations and copy number alterations Recurrent aberrations Gains within chromosome 8 (o) Multiple deletions (including in 14q) (c, p) 20q11.21 and 12p13.31(i, m, q, r, s) Small chromosomal aberrations occur both in mouse and human ESCs, but no syntenic recurrent aberrations have been identified. Most CNVs arise from selection for rare populations in the parental cells during reprogramming or culturing (u, v, w) The total number of CNVs decreases in culture(t) Specific genes have been suggested, such as BCL2L1 (i) and NANOG (m>, n) May be associated with pluripotency pseudo-genes, cancer-related genes, and genes within common fragile sites (m, o, t) Defects in DNA damage response and replication stress 
Frequency of aberrations Average of 109 CNVs per hiPSC line and 55 CNVs per hESC line (t) 10–25% of hESCs display the recurrent amplification of 20q11.21 (i, m) 13% of hESCs display the recurrent amplification of 12p13.31 (m) 
Single nucleotide variations (SNVs) Recurrent aberrations Not identified Not identified One study identified the same variants in four mouse iPS clones. This has not been observed in human PSCs. (w) Most SNVs can be traced back to the parental cells Shared SNVs were not observed between different iPS cell lines derived from the same somatic fibroblasts Replication defects 
Frequency of aberrations ∼11 point mutations in coding regions per clone (w) ∼6 point mutations in coding regions per clone (v, x, y) 
Aberration type Mouse PSCs Human PSCs Comparison of human and mouse PSCs Origin Gene enrichment Likely mechanism of formation 
Chromosomal aberrations Recurrent aberrations Gains 8 and 11Deletions 10qB and 14q(a, b, c, d, e, f) Gains 1, 12, 17, 20 and X (g, h, i, j, k, l, m, n) Human chromosome 17 is completely syntenic to the distal half of mouse chromosome 11(c) Most aberrations arise in culture during propagation (culture adaptation) (i, n) Difficult to analyze, as aberrations contain multiple genes Defects in chromosomal segregation during cell division 
Frequency of aberrations ∼38% in mESCs ∼23% in miPSCs (c) ∼25% of the aberrations involve chromosomes 8 and 11 (c) ∼32–34% in hESCs (i, n) ∼20% in hiPSCs (n) ∼50% of the aberrations involve chromosomes 1,12,17 or 20 (i) 
Subchromosomal aberrations and copy number alterations Recurrent aberrations Gains within chromosome 8 (o) Multiple deletions (including in 14q) (c, p) 20q11.21 and 12p13.31(i, m, q, r, s) Small chromosomal aberrations occur both in mouse and human ESCs, but no syntenic recurrent aberrations have been identified. Most CNVs arise from selection for rare populations in the parental cells during reprogramming or culturing (u, v, w) The total number of CNVs decreases in culture(t) Specific genes have been suggested, such as BCL2L1 (i) and NANOG (m>, n) May be associated with pluripotency pseudo-genes, cancer-related genes, and genes within common fragile sites (m, o, t) Defects in DNA damage response and replication stress 
Frequency of aberrations Average of 109 CNVs per hiPSC line and 55 CNVs per hESC line (t) 10–25% of hESCs display the recurrent amplification of 20q11.21 (i, m) 13% of hESCs display the recurrent amplification of 12p13.31 (m) 
Single nucleotide variations (SNVs) Recurrent aberrations Not identified Not identified One study identified the same variants in four mouse iPS clones. This has not been observed in human PSCs. (w) Most SNVs can be traced back to the parental cells Shared SNVs were not observed between different iPS cell lines derived from the same somatic fibroblasts Replication defects 
Frequency of aberrations ∼11 point mutations in coding regions per clone (w) ∼6 point mutations in coding regions per clone (v, x, y) 
a

Liu et al., 1997; bBrimble et al., 2004; cBen-David and Benvenisty, 2012b; dLiang et al., 2008; eSugawara et al., 2006; fSommer et al., 2010; gBen-David et al., 2011; hTaapken et al., 2011; iAmps et al., 2011; jDraper et al., 2004; kBaker et al., 2007; lMartins-Taylo et al., 2011; mLaurent et al., 2011; nMayshar et al., 2010; oPasi et al., 2011; pArlt et al., 2012; qNärvä et al., 2010; rLefort et al., 2008; sWerbowetski-Ogilvie et al., 2009; tHussein et al., 2011; uAbyzov et al., 2012; vGore et al., 2011; wYoung et al., 2012; xCheng et al., 2012; yRuiz et al., 2013.

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