Table 3.

Mitochondria, ER, MAM, and other localization of mTORC2

Pathway Mechanism of localization Cell type Technique Reference 
mTORC2 mTORC2 (mTOR, rictor) is mainly at the ER MDA-MB-435; A549 Confocal imaging and subcellular fractionation; PFA fixation; endogenous mTOR and rictor Boulbés et al., 2011  
mTORC1/2 mTOR is associated with mitochondria NIH3T3 Confocal imaging, subcellular fractionation, and immuno-EM; PFA fixation; endogenous mTOR Desai et al., 2002  
mTORC2 mTORC2 (mTOR; Sin1; rictor) is at MAM HeLa; MEF; mouse liver Confocal imaging, subcellular fractionation, and immuno-EM; PFA fixation; endogenous mTOR, Sin1, and rictor; validation of antibody Betz et al., 2013; Poston et al., 2013  
mTORC2 A part of mTORC2 (mTOR; rictor; Sin1) is nuclear; rapamycin treatment induces translocation of mTORC2 to the cytoplasm HEK293; IMR-90; NIH3T3 Subcellular fractionation Rosner et al., 2007; Rosner and Hengstschläger, 2008, 2011, 2012  
Yeast TORC2 TORC2 (TOR2; Bit61; Avo1-3) is at or near the plasma membrane, possibly cortical ER S. cerevisiae Subcellular fractionation, immuno-EM, live-cell imaging, and wide-field and confocal imaging; PFA fixation; TOR2-GFP, Bit61-GFP, Avo1-3-GFP Kunz et al., 2000; Wedaman et al., 2003; Aronova et al., 2007; Sturgill et al., 2008; Berchtold and Walther, 2009  
Pathway Mechanism of localization Cell type Technique Reference 
mTORC2 mTORC2 (mTOR, rictor) is mainly at the ER MDA-MB-435; A549 Confocal imaging and subcellular fractionation; PFA fixation; endogenous mTOR and rictor Boulbés et al., 2011  
mTORC1/2 mTOR is associated with mitochondria NIH3T3 Confocal imaging, subcellular fractionation, and immuno-EM; PFA fixation; endogenous mTOR Desai et al., 2002  
mTORC2 mTORC2 (mTOR; Sin1; rictor) is at MAM HeLa; MEF; mouse liver Confocal imaging, subcellular fractionation, and immuno-EM; PFA fixation; endogenous mTOR, Sin1, and rictor; validation of antibody Betz et al., 2013; Poston et al., 2013  
mTORC2 A part of mTORC2 (mTOR; rictor; Sin1) is nuclear; rapamycin treatment induces translocation of mTORC2 to the cytoplasm HEK293; IMR-90; NIH3T3 Subcellular fractionation Rosner et al., 2007; Rosner and Hengstschläger, 2008, 2011, 2012  
Yeast TORC2 TORC2 (TOR2; Bit61; Avo1-3) is at or near the plasma membrane, possibly cortical ER S. cerevisiae Subcellular fractionation, immuno-EM, live-cell imaging, and wide-field and confocal imaging; PFA fixation; TOR2-GFP, Bit61-GFP, Avo1-3-GFP Kunz et al., 2000; Wedaman et al., 2003; Aronova et al., 2007; Sturgill et al., 2008; Berchtold and Walther, 2009  

This table groups the most important reports that investigate TORC2 and mTORC2 localization. mTORC2 has consistently been identified in proximity to mitochondria, the ER, and MAM. Other studies suggest that mTORC2 is nuclear or at the plasma membrane. EM, electron microscopy; PFA, paraformaldehyde.

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