Table I.

Key targets identified from isolation of CLIC-positive fraction

Target Location Link to endocytosis 
4F2 cell surface antigen heavy chain/CD98 PM Receptor for Galectin3 and Integrin β-1 endocytosis 
78-kD glucose-regulated protein PM/ER Associates with lipid rafts and receptor for MHCI 
Actin, cytoplasmic 1 Cytosol Drives endocytic carrier formation 
Alpha-2-HS-glycoprotein/Fetuin PM/E Endocytosis mediated by AnnexinA2 
Annexin A2 Binds glycoproteins during endocytosis 
CD109 antigen PM Cell surface GPI-AP 
CD44 antigen PM Hyaluronan receptor 
Collagen ECM ECM constituent 
Fructose-bisphosphate aldolase A Cytosol Mediates Snx9-dynamin interaction 
Galectin-3 PM Mediates Integrin β-1 endocytosis 
Guanine nucleotide-binding protein Gα(i,o) PM Involved in STAT3 signaling 
Integrin β-1 PM Receptor for ECM 
Lysosome-associated membrane glycoprotein 2 Lysosomal membrane regulator 
Membrane-associated progesterone receptor component 1 PM/ER Involved in cholesterol homeostasis, rapidly cleared from the PM by endocytosis 
Moesin PM Links CD44 to cortical actin to drive features in the PM 
Myoferlin (Fer-1–like protein 3) PM Involved in Ca2+-promoted membrane fusion; binds to AHNAK 
Neuroblast differentiation-associated protein AHNAK PM Binds Myoferlin and Dysferlin and with Annexin2 mediates membrane fusion 
Prolow-density lipoprotein receptor-related protein 1 (LRP1) PM Contains both tyrosine and di-leucine sorting motifs; binds saposin on cell surface 
Prosaposin PM/E Co-receptor with LRP1; binds GM1 
Rac1 PM/E Induces membrane ruffling 
Rab-11A Endosome recycling 
Rab-14 Endosome recycling 
Rab-7a Late endosome fusion and motility 
Rab-5A Early endosome fusion and motility 
Rap-1b PM Integrin β-1 endocytosis and focal adhesion turnover; mediates STAT3 
Receptor expression-enhancing protein 5 PM Involved in Rab-mediated membrane trafficking 
Reticulon-4 PM/ER Tubulates membranes; binds NogoR, a GPI-AP at the PM 
Thy-1 membrane glycoprotein PM GPI-AP; regulates FAK signaling 
Tmp21 PM Contains KDEL sequence, traffics to the PM independently from p24 
Vimentin Cytosol With Annexin A2 regulates FAK signaling 
Target Location Link to endocytosis 
4F2 cell surface antigen heavy chain/CD98 PM Receptor for Galectin3 and Integrin β-1 endocytosis 
78-kD glucose-regulated protein PM/ER Associates with lipid rafts and receptor for MHCI 
Actin, cytoplasmic 1 Cytosol Drives endocytic carrier formation 
Alpha-2-HS-glycoprotein/Fetuin PM/E Endocytosis mediated by AnnexinA2 
Annexin A2 Binds glycoproteins during endocytosis 
CD109 antigen PM Cell surface GPI-AP 
CD44 antigen PM Hyaluronan receptor 
Collagen ECM ECM constituent 
Fructose-bisphosphate aldolase A Cytosol Mediates Snx9-dynamin interaction 
Galectin-3 PM Mediates Integrin β-1 endocytosis 
Guanine nucleotide-binding protein Gα(i,o) PM Involved in STAT3 signaling 
Integrin β-1 PM Receptor for ECM 
Lysosome-associated membrane glycoprotein 2 Lysosomal membrane regulator 
Membrane-associated progesterone receptor component 1 PM/ER Involved in cholesterol homeostasis, rapidly cleared from the PM by endocytosis 
Moesin PM Links CD44 to cortical actin to drive features in the PM 
Myoferlin (Fer-1–like protein 3) PM Involved in Ca2+-promoted membrane fusion; binds to AHNAK 
Neuroblast differentiation-associated protein AHNAK PM Binds Myoferlin and Dysferlin and with Annexin2 mediates membrane fusion 
Prolow-density lipoprotein receptor-related protein 1 (LRP1) PM Contains both tyrosine and di-leucine sorting motifs; binds saposin on cell surface 
Prosaposin PM/E Co-receptor with LRP1; binds GM1 
Rac1 PM/E Induces membrane ruffling 
Rab-11A Endosome recycling 
Rab-14 Endosome recycling 
Rab-7a Late endosome fusion and motility 
Rab-5A Early endosome fusion and motility 
Rap-1b PM Integrin β-1 endocytosis and focal adhesion turnover; mediates STAT3 
Receptor expression-enhancing protein 5 PM Involved in Rab-mediated membrane trafficking 
Reticulon-4 PM/ER Tubulates membranes; binds NogoR, a GPI-AP at the PM 
Thy-1 membrane glycoprotein PM GPI-AP; regulates FAK signaling 
Tmp21 PM Contains KDEL sequence, traffics to the PM independently from p24 
Vimentin Cytosol With Annexin A2 regulates FAK signaling 

E, Endosomes; ECM, extracellular matrix; ER, endoplasmic reticulum; PM, plasma membrane.

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