Table 2.

NK cell activation and metabolic modulation in response to different stimuli

StimuliDosageNK cell responseSource of NK cellsReference
High-dose or low-dose IL-12 and IL-18 IL-12 (1 ng/ml), IL-18 (1 ng/ml); or IL-12 (10 ng/ml), IL-18 (50 ng/ml) Activation and IFN-γ production independent of glycolysis or OXPHOS Splenocytes (murine) Keppel et al., 2015  
Anti-NK1.1 or anti-Ly49D 20 μg/ml Activation and IFN-γ production dependent on glycolysis and OXPHOS Splenocytes (murine) Keppel et al., 2015  
High-dose IL-15 and anti-NK1.1 or anti-Ly49D IL-15 (100 ng/ml), anti-NK1.1, or anti-Ly49D (20 μg/ml) Activation and IFN-γ production independent of glycolysis or OXPHOS Splenocytes (murine) Keppel et al., 2015  
High-dose IL-15 IL-15 (100 ng/ml) or IL-15/IL-15Rα complex (50 or 100 ng/ml) Activation, IFN-γ production, and increased metabolic profile (glycolysis and OXPHOS) dependent on mTOR signaling Splenocytes (murine) Marçais et al., 2014; Nandagopal et al., 2014  
Low-dose IL-15 IL-15 (10 ng/ml) or IL-15/IL-15Rα complex (10 ng/ml) Cell survival and viability dependent on STAT5 phosphorylation Splenocytes (murine) Marçais et al., 2014; Nandagopal et al., 2014  
IL-2 and IL-12 IL-2 (20 ng/ml), IL-12 (10 ng/ml) Activation and IFN-γ production; increased metabolic profile (glycolysis and OXPHOS) independent of glutamine-fueled TCA cycle; increased cMyc expression dependent on glutamine availability and SLC7A5 activity Splenocytes (murine) Loftus et al., 2018  
IL-12, IL-15, and IL-18 (CIML NK cells) IL-12 (10 ng/ml), IL-15 (10 ng/ml), IL-18 (50 ng/ml) More IFN-γ production after re-stimulated by IL-12 (10 ng/ml) and IL-15 (100 ng/ml) or anti-Ly49H or anti-NK1.1 (5 μg/ml) Splenocytes (murine) Cooper et al., 2009  
IL-12 (10 ng/ml), IL-15 (100 ng/ml), IL-18 (50 ng/ml) More IFN-γ production after re-stimulated by IL-12/15/18 (10, 100, 50 ng/ml, respectively) or K562 cells; shift towards glycolysis upon activation with short-term increased OXPHOS; increased expression of nutrient transporters (CD98 and GLUT1) PBMC (human) Terrén et al., 2021  
IL-12 and IL-15 IL-12 (30 ng/ml), IL-15 (100 ng/ml) Activation and IFN-γ production dependent on elevated OXPHOS; upregulated glycolysis and OXPHOS insensitive to rapamycin PBMC (human) Keating et al., 2016  
IL-2 500 U/ml Activation and IFN-γ production dependent on elevated OXPHOS; upregulated glycolysis and OXPHOS dependent on mTORC1 PBMC (human) Keating et al., 2016  
IL-2 and anti-NKG2D IL-2 (200 U/ml), anti-NKG2D (5 μg/ml) Activation and IFN-γ production; upregulated expression of SLC1A5 and CD98 mediated by mTORC1 (a prerequisite of following NKG2D-mediated activation) PBMC (human) Jensen et al., 2017  
StimuliDosageNK cell responseSource of NK cellsReference
High-dose or low-dose IL-12 and IL-18 IL-12 (1 ng/ml), IL-18 (1 ng/ml); or IL-12 (10 ng/ml), IL-18 (50 ng/ml) Activation and IFN-γ production independent of glycolysis or OXPHOS Splenocytes (murine) Keppel et al., 2015  
Anti-NK1.1 or anti-Ly49D 20 μg/ml Activation and IFN-γ production dependent on glycolysis and OXPHOS Splenocytes (murine) Keppel et al., 2015  
High-dose IL-15 and anti-NK1.1 or anti-Ly49D IL-15 (100 ng/ml), anti-NK1.1, or anti-Ly49D (20 μg/ml) Activation and IFN-γ production independent of glycolysis or OXPHOS Splenocytes (murine) Keppel et al., 2015  
High-dose IL-15 IL-15 (100 ng/ml) or IL-15/IL-15Rα complex (50 or 100 ng/ml) Activation, IFN-γ production, and increased metabolic profile (glycolysis and OXPHOS) dependent on mTOR signaling Splenocytes (murine) Marçais et al., 2014; Nandagopal et al., 2014  
Low-dose IL-15 IL-15 (10 ng/ml) or IL-15/IL-15Rα complex (10 ng/ml) Cell survival and viability dependent on STAT5 phosphorylation Splenocytes (murine) Marçais et al., 2014; Nandagopal et al., 2014  
IL-2 and IL-12 IL-2 (20 ng/ml), IL-12 (10 ng/ml) Activation and IFN-γ production; increased metabolic profile (glycolysis and OXPHOS) independent of glutamine-fueled TCA cycle; increased cMyc expression dependent on glutamine availability and SLC7A5 activity Splenocytes (murine) Loftus et al., 2018  
IL-12, IL-15, and IL-18 (CIML NK cells) IL-12 (10 ng/ml), IL-15 (10 ng/ml), IL-18 (50 ng/ml) More IFN-γ production after re-stimulated by IL-12 (10 ng/ml) and IL-15 (100 ng/ml) or anti-Ly49H or anti-NK1.1 (5 μg/ml) Splenocytes (murine) Cooper et al., 2009  
IL-12 (10 ng/ml), IL-15 (100 ng/ml), IL-18 (50 ng/ml) More IFN-γ production after re-stimulated by IL-12/15/18 (10, 100, 50 ng/ml, respectively) or K562 cells; shift towards glycolysis upon activation with short-term increased OXPHOS; increased expression of nutrient transporters (CD98 and GLUT1) PBMC (human) Terrén et al., 2021  
IL-12 and IL-15 IL-12 (30 ng/ml), IL-15 (100 ng/ml) Activation and IFN-γ production dependent on elevated OXPHOS; upregulated glycolysis and OXPHOS insensitive to rapamycin PBMC (human) Keating et al., 2016  
IL-2 500 U/ml Activation and IFN-γ production dependent on elevated OXPHOS; upregulated glycolysis and OXPHOS dependent on mTORC1 PBMC (human) Keating et al., 2016  
IL-2 and anti-NKG2D IL-2 (200 U/ml), anti-NKG2D (5 μg/ml) Activation and IFN-γ production; upregulated expression of SLC1A5 and CD98 mediated by mTORC1 (a prerequisite of following NKG2D-mediated activation) PBMC (human) Jensen et al., 2017  

CIML, cytokine-induced memory-like; SLC7A5, solute carrier family 7 member 5; SLC1A5, solute carrier family 1 member 5.

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